This draft has only been discussed superficially




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84  Polemoniaceae (Draft: R. Elven)

THIS DRAFT HAS ONLY BEEN DISCUSSED SUPERFICIALLY

84.1  Phlox L. (1753), Sp. Pl. 151.
The Phlox sibirica aggregate (P. alaskensis, P. richardsonii, P. sibirica)

Comments:

(1) The three entities included are obviously closely related and are treated as subspecies by Hultén (1968) and Löve & Löve (1975). The two American plants ('alaskensis', 'richardsonii') are fairly well separated morphologically and as far as I know without intermediates. Hultén (1968) was of the opinion that the Siberian ('sibirica') and Alaskan ('alaskensis') plants were the same but that 'richardsonii' deserved status as a separate subspecies. Cody (1996) reverted to three species. See the discussion in Hultén (1968, Ark. Bot. p. 110-111). Until further comments all three are entered as species but with 'warnings'. Opinions? (Elven)
84.1.1  Phlox sibirica L. (1753), Sp. Pl. 153.

S P. sibirica L. subsp. sibirica

T [Described from N Siberia.]

2n= 14 (2x).

2nD Zhukova (1967a).

G SIB RFE

Comments:
84.1.2  Phlox alaskensis Jordal (1952), Rhodora 54: 38.

S P. richardsonii Hook. taxon alaskensis (Jordal) Wherry (1955), Morris Arb. Monogr. 3: 217; P. sibirica L. subsp. alaskensis (Jordal) Á. Löve & D. Löve (1976), Bot. Not. 128: 518.

T [Described from N Alaska: Wiseman, leg. Jordal 2198.]

2n= 14 (2x).

2nD Johnson & Packer (1968 Ala, as P. sibirica); Mulligan & Porsild in Löve (1970a).

G RFE ALA CAN

Comments:

WARNING! Should probably be merged with P. sibirica without special rank.
84.1.3  Phlox richardsonii Hook. (1840), Fl. Bor. Amer. 2: 73, t. 160.

S P. sibirica L. subsp. richardsonii (Hook.) Hultén (1968), Ark. Bot., ser. 2, 7, 1: 111.

2n=

2nD


G ALA CAN

Comments:



WARNING! Should probably be reduced to a subspecies of P. sibirica.
84.1.4  Phlox hoodii Richardson in Franklin (1823), Bot. App., ed. 2, 6, t. 28.

S

2n= 28 (4x).



2nD Flory (1937); Crawford in Löve (1971a).

G ALA CAN

Comments:

84.2  Collomia Nutt. (1818), Gen. N. Amer. Pl. 1: 126.
84.2.1  Collomia linearis Nutt. (1818), Gen. N. Amer. Pl. 1: 126.

S

2n=



2nD

G ALA*? CAN*?

Comments:

(1) Approaches the Arctic in Alaska and reaches it in the Hudson Bay region. It may, however, only be casual in the northernmost sites and therefore to be excluded. (Elven)



WARNING! Genus and species might be excluded as not stable in the Arctic.

84.3  Polemonium L. (1753), Sp. Pl. 162.

Comments:

(1) Löve & Löve (1975) include an additional species as arctic: P. villosum J. Rudolph ex Georgi (1800), Beschr. Russ. Reich. 3, 4: 771, with subsp. villosum (Siberia, Far East) and subsp. pacificum (V. Vassil.) Á. Löve & D. Löve *** (Siberia). According to Tzvelev in Tolmachev & Yurtsev (1980, Fl. Arct. URSS 8 p. 220-221), P. villosum is a nomen ambiguum and is placed in the synonymy of P. boreale. What reality has this 'entity' and does it reach the Arctic as defined? (Elven)
84.3.1  Polemonium caeruleum L. (1753), Sp. Pl. 162.

S

2n= 18 (2x).



2nD Löve & Löve (1975) list numerous counts, two as arctic.

G NOR RUS SIB

Comments:

(1) In North Norway this species is coastal and native and reaches the Arctic in at least one site (in Lebesby) in established sand dunes. Hultén & Fries (1986) map it from Polar Ural, lower Ob and lower Jenisei, within the Arctic as defined and Tzvelev lists it from Malozemelskaya Tundra. (Elven)


84.3.2  Polemonium acutiflorum Willd. in Roem. & Schult. (1819), Syst. Veg. 4: 792.

S P. campanulatum (Th. Fr.) H. Lindb. ex Lindm. (1918), Sv. Fanerogamfl. 457.

2n= 18 (2x).

2nD Löve & Löve (1975) list numerous counts, several as arctic.

G RUS SIB RFE ALA CAN

Comments:

(1) Tzvelev in Tolmachev & Yurtsev (1980, Fl. Arct. URSS 8) separates between P. acutiflorum and P. campanulatum and gives the latter from, e.g., the Murman area (Kola Peninsula), Bolshezemelskaya Tundra, lower Ob, Gydan Tundra, lower Jenisei, Kolyma, and Chukchi Peninsula, all of these areas generally within what he also maps for P. acutiflorum. He also synonymises P. villosa subsp. pacifica (see comment above) with P. campanulata. I can see no good reason for this division yet and have synonymised the two. (Elven)

(2) Even if P. acutiflorum closely approaches the Arctic also in North Norway, it stops abruptly about 15-25 km from our defined boundary in several places. (Elven)


84.3.3  Polemonium boreale Adams (1817), Mém. Soc. Nat. Moscou 5: 92.

S ?P. hyperboreum Tolm. (1927), Feddes Repert. 23, 18-25: 273; ?P. boreale Adams subsp. hyperboreum (Tolm.) Á. Löve & D. Löve ***

2n= 18 (2x).

2nD Löve & Löve (1975) list several counts, most as arctic, for their subsp. boreale.

G NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) The 'hyperboreum' taxon is synonymised with P. boreale by Tzvelev (Fl. Arct. URSS 8) but separated as a subsp. hyperboreum of P. pulchellum by Löve & Löve (1975) and as a separate species by Hultén (1968). Opinions?

Löve & Löve (1975) accept two subspecies of P. boreale (excluding 'hyperboreum'); the second is subsp. humile (Willd.) Á. Löve & D. Löve (1976), Bot. Not. 128: 517 [basionym: P. humile Willd. ex Roem. & Schult. (1819), Syst. Veg. 4: 792, non Salisb. (1796), Prodr. 125], said to be "the northern Siberian race of the species", and also given with 2n=18 (Matveeva & Tikhonova in Bolkhovskikh et al. 1969). (Elven)


84.3.4  Polemonium pulcherrimum Hook. (1830), Curtis' Bot. Mag., n. s., 4: t. 2979.

S P. pulcherrimum Hook. subsp. pulcherrimum

2n= 18 (2x).

2nD Löve & Löve (1975) list five counts, one as arctic.

G ALA CAN

Comments: See P. boreale.



85  Hydrophyllaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED

85.1  Phacelia Juss. (1789), Gen. 129.
85.1.1  Phacelia franklinii (R. Br.) A. Gray (1856), Manual, ed. 2, 329.

B Eutaca franklinii R. Br. in Frankl. (1823), Bot. App. t. 27.

S

2n=


2nD

G CAN


Comments:

(1) Eutaca franklinii is also said to have been first published by R. Br. (1837), Trans. Linn. Soc. 17: 277. (Elven)



(2) Occurs in the Arctic at least at Hudson Bay (north to Churchill, Scoggan 1978) and also reaches far north in Mackenzie District. A border case. (Elven)


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