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vers. 2008

Provisional key for the genus


MOLLISIA s. l.

(with provisional inclusion of some of the species created by Svrcek (SV) and LeGal (LG), which had to be keyed out KOH+ as well as KOH- because of missing descriptions. They all have not been examined by me)



Characteristics of the genus:

Continuous refractive vakuole in the paraphyses (KOH-soluble!), as well as asci with  slender, tapering base. Concerning the marginal cells and the excipulum (textura globulosa vs. angularis) the limitation versus Pyrenopeziza is sometimes difficulte and on working with exsiccates classification is in many cases only tentatively practible, because the refractive vacuoles are only visible in fresh material.



Abbreviations:
IKI Iod-Kalium-Iodid (Lugols solution)

- negative

bb blue

rr red


rb red, but in lower concentrations of Iod blue

It is very important to prove the ascus-reaction without KOH-pretreatment in a H2O-preparation, because the red reaction of the porus will be blue together with KOH!
Oil total content of oil-drops in the spore

0 without oil-content

1 ca. 3% of the volume of the spore consists of oil-drops

2 ca. 10% ------------------------- " --------------------------

3 ca. 25% ------------------------- " --------------------------

4 ca. 50% ------------------------- " --------------------------

5 ca. 80% ------------------------- " --------------------------
KOH yellow or violet reaction

reaction of the content of the paraphyses when getting in contact with KOH 3%. Reaction may be weak or strong, only for seconds or permanent. Weak reactions are usualy only be observed with fresh material, but the strong reactions (e. g. M. rosae) are sometimes even in 130 year old specimens still clearly visible. When working with exsiccates no reaction with KOH is not a clear statement, but if the reaction is positive (yellow or violet) it is always a good feature.


Unless stated otherwise, all details and measurements are from living collections observed in water. Exsiccates were studied in KOH 3% except for proving the ascus-reaction.
Names underlined: Own collections have been made, or at least own revisions of exsiccates.

1a On wood, bark 2

1b On remnants of herbs (incl. Rubus) 50

1c On remnants of grasses s. l. 70

1d On leaves of trees or shrubs (leaves of herbs and grasses: 1b or 1c) 120

1e On mosses 140




WOOD, BARK
2a Ascus reaction rr 3

2b Ascus reaction bb 5

2c Ascus reaction negative 45

2d Ascus reaction rb, margin cells conspicious, 35-100 µm, hyaline, KOH negative, Sp. 8-


14 x 2,8-3,5 µm, underneath rotten Quercus logs in winter Mollisia elegantior

M. elegantior Baral comb. nov. ined.


3a Sp. 7-10 x 2,5-3,2 µm, growing ± submerged, marginal cells inconspicuous

M. "dextrinoidea"

3b Sp. 9-12 x 2,2-3 µm, not submerged M. „Malaval“


5a Subhymenium (text. intr.) made of brownish and loosely arranged hyphae (at least in
mature apo., needs sometimes several preparations), sp. 6-13 x 2,5-3,3 µm, oil 0(+),
KOH- M. lividofusca

According to LeGal & Mangenot, M. fallax could be distinguished by flat, discinoid apothecia, smooth excipulum and shorter spores with oil drops.

When spores 20-28 µm long  40**

5b Subhymenium built of hyaline hyphae 7


7a Marginal cells very conspicuous, evidently > 50 µm, blackish-brown and mostly
encrusted and KOH-reaction always strongly lemon-yellow 8

7b Marginal cells less conspicuous, KOH-reaction yellow or negative 10


8a Sp. 6-9 x 2-2,8 µm, oil 2(+), Asci without croziers, on Rosa spec. (+ Rubus?) M. rosae

8b Sp. 9-14 x 2,2-3 µm, oil 3-4, Asci with croziers, on Prunus spinosa M. prunicola

8c Sp. 6-12,5 x 1,8-2,5 µm, oil 0, Asci with croziers, Apo. up to 7 mm , semihypogaeic on
rotten Quercus-truncs (often together with Hymenochaete rubiginosa, Dasyscyphella
nivea
) „Haglundia“ perelegans

M. perelegans Baral comb. nov. ined.
10a KOH-reaction ± distinctly yellow 11

10b KOH-reaction negative 20

LG KOH unknown, spores (8)10-23,5 x 1,5-2,5 µm, oil unknown, probably appr. 2, medulla
built of parallel, densly woven hyphae M. caesia
11a Sp. septate already in the asci, 14-17 x 2,8-3 µm, ascus of Pyrenopeziza-Typ, 65 x 8 µm

Pyrenopezizacoriariae

11b Sp. not septate within mature asci, of other size 12
12a Oil 2-3(4) (compare also 7 u. 7*) 15

12b Oil 0-1 17

12c Oil either 1 (many very tiny droplets) or 1,5-2 (several bigger drops), if 1 then ascus
reaction strong bb, if 1,5-2 than ascus reaction weakly bb, apo. shortly (blackish)
stipitate M. perparvula

Two specimens in H (PAK 2922, 2930) are identical according to Huhtinen, but seem to be distinct to me.


15a Sp. 13-18(20) x 3,5-4,2 µm, outside the ascus with 1-3 septae, marginal cells conspicious,
egg-shaped with somewhat pointed apex M. ulicis (ss. Priou)

15b Sp. more narrow and shorter, even when mature with max. 1 septum 16


16a Sp. (9?)10-15 x 2,2-3,2 µm, Q > 4 (in average), apo. ± patellate, cup-shaped, generally
blueish-grey, subicular hyphae up to 6 µm broad M. fusca

16b Sp. idem, but oil only 1-2, marginal cells greybrown and quite conspicuous, growing only


in spring, on Fagus-cupules (sometimes on small twigs), often together with Brunnipila
fuscescens
, Capitotricha fagiseda und Lachnum virgineum, M. "fagicola"

This is very probably M. faginea Velenovský.

SV Sp. 10-15 x 3,5-4,5 µm, oil 1-2: small drops accumulated in the polar ends of the spore,
marginal cells long, KOH perhaps positive M. sericeomarginata
17a Sp. 2-3 µm broad 18

17b Sp. > 3 µm broad 19


18a Sp. 6-8(9) x 2-2,8 µm, oil 0 M. "pyrenopezizoides"

On Rubus idaeus, without croziers  compare M. alcalireagens Svrcek

18b Sp. 8-12,5 x 2,2-3 µm, oil 0+, hymenium and margo white, very moistened slightly
greyish, on Prunus spinosa and Crataegus M. "albogrisea"

18c Sp. 7-10 x 1,5-2,2 µm, oil ca. 1, hymenium bluish-grey, with conspicuous blackish


margo, marginal cells conspicuous, conicol on Pinus sylvestris M. „neffii“

18d Sp. 7,5-10 x 2,2-2,8 µm, oil 1, apo. small, watery greyish, ash-grey, without clearly


visible margo, textura ± angularis (to be checked!), exc. not up to margo brownish, on
Picea, Larix-cones, Prunus M. "conifericola"

Identical seems to be PAK 4398: M. leucostigma (with „?“), but spores 11-13 x 2-2,2 µm (already germinating) and margo more conspicuous. Fuckels N. leucostigma has totally different spores and a darker margo!


19a Sp. 9-12 x 3-4 µm, Q 2,6-3,8, apo. cushion-like, chalk-white, sometimes with a slight
greyish hue, without subiculum, always submerged M. “pulviniformis”

The original diagnosis of M. uda by Persoon is obviousely quite different from this species which therefore needs a new name. According to M. Nauta (in litt.) no authentic material is left in L.

19b Sp. 8-14 x 3,3-4,2 µm, amyloid-ring (porus) T-shaped, ect. Exc. very thick (according to
Baral) M. sericeomarginata

SV KOH yellow?, Sp. 10-15 x 3,5-4,5 µm, oil 1 (and more?), Apo. not submerged



M. sericeomarginata
20a Sp. already within the vital asci with 1 or more septae 21

20b Dead spores usually with septae, oil >2,5 23

20c Dead and vital spores usually not septate (or rarely before germinating) 25
21a Sp. already in the ascus 1-3-septated, 10-16 x 3,2-4,5 µm, oil 0, in IKI with weakly red
sheath (?), on thick blackish subiculum N. "karkanoszeae"

further collections by H. O. Baral from Spain and Thüringen are very likely identical.

21b Sp. in the asci always with 1 septum, without blackish subiculum 22
22a Sp. 10-16 x 3-3,5 µm, oil 0,5-1, apo. conglomerated, corticolous M. „septispora“

22b Sp. (7)8-10(11) x 2,5-3 µm, oil 0-0,5 (tiny droplets along the septum), apo. gregarious,


lignicolous M. „atlantica
23a Sp. 33-56 x 2,5-4 µm, 4-celled, oil 2-3, hymenium opalescent blueishgrey, on Calluna-
roots at the base of dying plants, often nearly in the earth, also on Ericaceae

M. (“Belonopsis”) obscura

23b Sp. 11-18 x 2,5-3,5, 0-2-4-celled, oil 4-5, hymenium olive-greenish, drying yellowish (like


mustard), growing only in very wet places, sometimes submers M. ventosa
25a Cells of the ect. exc. only at the apo. base brownish (apo. adhered  punctiform ) 26

25b Cells of the ect. exc. +/- up to the margin brownish coloured 28


26a Apo. watery-grey, breaking through the cortex in fascicles, sp. 8-12(-14 nach Declerq) x
2-3, oil 2 (always?) M. benesuada

Many quite similar specimens have been observed and the problem in delimitation of the species seem to be following two points: Is the ect. exc. always (sub)hyaline and must M. benesuada grow fasciculated?

This species is unclear to me!

26b Apo. amber-coloured, singly growing, sp. 8,5-11 x 2-2,5, oil 0, IKI bb 1(-2), on Rubus


idaeus
M. "amberina"

26c Sp. 5-9(10) x 1,5-2,5 µm, cells of the ect. Exc. only ochreous, often some coloured cells


up to the margo, exclusive on Alnus-cones M. amenticola
27a Oil 0(+) 30

27b Oil appr. 2 35

27c Oil >3-4+ 40

GL Oil unknown, probably appr. 2, sp. (8)10-23,5 x 1,5-2,5 µm, medulla built of parallel,


dense hyphae, KOH unknown M. caesia

GL Oil unknown, probably <1, sp. 6-10(11) x 1-1,7 µm M. cinereoolivascens


30a Sp. 5-9(10) x 1,5-2,5 µm, ekt. exc. only ochreous, on Alnus-cones M. amenticola

30b Ect. Exc. darker brown 31


31a On cones of conifers, Sp. 6-14 x 2-3,2 µm, subhymenium always hyalin M. strobilicola

doubtful species and never found by me! All collections labelled M. strobilicola have been found to be identical with M. lividofusca ( 5), because of the coloured subhymenium.

31b Host otherwise 32
32a Sp. usually not exceeding 10 µm, at least in average 33

32b Sp. usually not < 10 µm, at least in average 34


33a Sp. 7-11(12?) x 2,5-3 µm, on wood, marginal cells not conspicuous M. cinerea

33b Sp. 7,5-9 x 1,8-2 µm on Rubus-cains (exclusively?), marginal cells long and con-


spicuous, with clavate last cell M. clavata

33c Sp. 7-10,5 x 1,8-2,2 µm, marginal cells with up to 40 µm long, +/- hyaline end cell, on


wood M. ladae

SV Sp. 5,5-10 x 2 µm [NH4OH], oil 0, excipulum with olive-greenish colour (al least in


NH4OH) M. olivaceocinerea

Coll. JPP9851 (Priou) keyed out here and belongs probably to the M. atrata complex.

GL Sp. 6-10(11) x 1-1,7 µm, oil unknown, probably <1 M. cinereoolivascens

GL Sp. 6-8,7-10(12) x 1,5-2 µm (exs., in BWB). KOH unknown M. cinerella Sacc.

GL Sp. (5)6-8-10(11) x 1,7-2,2 µm (exs. in BWB). KOH unknown M. undulatodepressula
34a On Picea abies, sp. 9-14 x 2,5-3 µm, hymenium blueish-grey, with strong blackish
subiculum M. "subcinerella"

several collections made by Lagerström in H sub M. melaleucoides could be identical. Sp. 8,5-12 x 2-3 µm, oil 0-0,5(1), marginal cells hyaline at the margo and very long (up to 60 µm), below the Margo vesicular, brownish. M. melaleucoides ss. Karsten is M. lividofusca.

34b Sp. 12-16 x 2,5-3 µm, oil 0-0,5, marginal hairs long and claviform, hence the margo
delicately ciliate (KOH perhaps +, collection overmature) M. villosa

Fund JPP 9852 (Priou) keys out here, but has a different textura (medullary exc. lacking) and reminds at a „Haglundia“.

34c Sp. 10-17 x 2-3,2 µm (dead), oil 1(+), marginal cells long club-shaped, sometimes 2-
celled, margin not ciliate M. velenovskyi

34d Sp. 9-11(13) x 2-3 µm (dead), oil 1-1,5(2?), marginal cells multicellular M. peruni

34e Sp. 9-12(13) x 2,5-3 (vital), 8,5-11 x 1,7-2,2 µm (dead), oil 0-1, some tiny doplets in each
pole, apo. whitish, light to blueish-grey, marginal cells inconspicous, margin usually well
develloped, submerged in fast floating clear water rivulets on decidous wood M. rivularis

SV On decorticated coniferous wood, sp. 7-12(15) x 3-3,5 µm [dead], oil 0 or with one small


droplet in each end, with blackish granulae in the hymenium when mounted in Melzers

M. ponticulorum
35a Sp. 9-16(19) x 2,5-3,2 µm, with two big drops in each end (+ sometimes few more
smaller drops, hymenium light ochraceous, never (?) grey or blackish, corticol (always?)

M. discolor

How to classify collections with spores only up to 11(12) µm ???

35b Sp. 8-12 x 2,8-3,5 µm, oil 2, with several not too big drops, hymenium whitish, at most
pale creme-coloured, mainly occuring in spring (?) M. melaleuca

35c Sp. 12-15 x 3, oil 3, germinating by conidia, on wood, with ± rich subiculum (according to


Baral) M. cinerella

35d Sp. 9-11(13) x 2-3 µm (dead), oil 1-1,5(2?), marginal cells multicellular M. peruni


40a Sp. 12-15 x 2,3-2,8 µm, oil 4, hymenium blueish-grey, on Vaccinium uliginosum
(according to Baral) M. vaccinii

40b Sp. 20-28 x 2,5-3,2 µm, oil 5, hymenium ochraceous, always (?) with cristals in the


medulla, subhymenium generally with ochraceous hyphae M. ramealis

40c Sp. 11-18 x 2,5-3,5 µm, oil 4-5, hymenium olivaceous-greenish, drying ochre-yellowish


(like mustard) sp. mature (only overmature?) often with 1(-3) septae M. ventosa
45a Sp. 35-49 x 2,2-3 µm, up to 8-celled, on Erica arborea M. ericae

45b Sp. shorter and without septae 46


46a Sp. 2-3 µm broad 47

46b Sp. 1,2-2 µm broad 48


47a Marginal cells (dark) brown, 2-4-celled, end cell  clavate, apo. drought-resistent, not
fasciculate, sp. 6-10 x 2-3 µm, oil 1(+), M. ligni

47b Marginal cells subhyalin, end cells nearly cylindrical, apo. fasciculate, sp. (7)8-10(12) x


2,2-3 µm . oil 1(-2) M. depressula

concept is based on M. depressula x inexula (PAK 4373). Uncertain which name is to be used.

SV Sp. 7-9 x 2,5-3 µm [living], generally septated, oil ca. 1 (biguttulate), globose cells yellow
coloured and this pigment soluble, apo. olive(yellowish-)greenish M. viridula

perhaps only an overmature collection of ... ???


48a Sp. 5-6,8 x 1,2-1,8 µm, apothecia fasciculate, on ?stromatised wood M. caespiticia

48b Sp. 3,5-5,5 x 1,2-1,8 µm (PAK 4406: 5-7,5 x 1,8-2,2, if identical?), apo. not fasciculate,


marginal cells  conspicuous, subhyaline, KOH - M. sublividula (PAK 4408)

in PAK (H) are 3 collections of M. sublividula, one of them is a Cistella spec., the other two are not clearly identical.



HERBS, FERNS
50a Oil >2 51

50b Oil 0-1 56


51a Sp. 11-13 x 1,5-2 µm, oil 2-3, consisting of small droplets, on Eupatorium (and Senecio
fuchsii
?) M. coerulans

51b Sp. < 11 µm 52

51c Sp. > 14 µm 54
52a KOH-reaction clearly yellow, sp. 8-11 x 1,8-2,5 (exs.), oil 2-3 M. ?polygoni

52b KOH-reaction negative, sp. 8-11 x 1,8-2,2 (exs.), oil 2-3, consisting of small droplets, on Polygonum M. polygoni

Fund WU 8580: Sp. 7-10 x 1,8-2,2 (exs.), oil 3+, consisting of few bigger, confluent drops.
54a Sp. 14-19 x 2-2,5(3) µm, oil 2-3, consisting of small and bigger drops, on Solidago
virgaurea
(similar to Pyrenopeziza chailletii, but with refractive vacuoles in the para-
physes) M. "solidaginis"

54b Sp. (17)20-23(26) x 3-4 µm, oil 2-3, consisting of small droplets, on Petasites petioles


(no refractive vacuoles!) Pyrenopeziza baraliana
56a Marginal cells inconspicuous, without remarkable features 57

56b Marginal cells somehow remarkable, conspicuous 60


57a Sp. 6-11 x 1,5-2,2 µm, oil 1(+), few small droplets in one ore both ends, KOH indistictly +
(dissolving very quick) M. revincta

57b Sp. same, oil 1(-), KOH absolute negativ (?), only on Filipendula (?) M. ?revincta

57 und 57* are very likely conspecific.

M. palustris Roberge is (according to the examination of the holotypus (in M) by H. O. Baral) perhaps only distinct from 57 and x or 57* because of KOH-negative reaction and growth restricted to grasses.

SV Reaction with NH4OH und KOH 3% strongly yellow, sp. 6-9 x 1,5-2 µm [living?], oil 0,5-1,


tiny droplets in one end (also in both?), ends of the spore rounded, on Rubus (only?)

M. alcalireagens

differences to M. „pyrenopezizoides“ ?


61a Sp. 8,5-11 x 2-2,5 µm, oil 0 (?), apo.  up to 3 mm, marginal cells often with extended
apex (constant feature?), KOH - M. adenostylidis

61b Sp. 7,5-9 x 1,8-2 µm, on Rubus-cains (exclusively?), marginal cells long and


conspicuous, with clavate last cell, paraphyses sublanceolate (always?) M. clavata

SV Sp. 5-8 x 1,3-1,5 µm, oil ca. 1: biguttulate, KOH ???, marginal cells hyalin,


conspicuousely long, 1-2-celled M. potentillae-erectae

most probably a Pyrenopeziza spec.!



GRASSES
70a Hymenium and medulla with lilac reaction when adding KOH, sp. 6-9 x 1,2-1,8 µm,
hymenium fresh orange, on Zea, Phyllistachis and Corylus M. „aurantioviolascens“

Only french collections. Same reaction as in Scutomollisia russea und purpurea, but no scutum and much smaller spores.

70b No lilac reaction when adding KOH 75
75a On Phragmites communis 76

75b On other grasses 85


76a Apo. with big masses of cristals in the ent. Exc. and the medulla 77

76b Apo. without cristals 80


77a Sp. (2-)4-celled, 26-31 x 3-4 µm (from exs.) M. mediella

Exs. Vestergren 1764 (= Trichobelonium distinguendum Sydow)

77b Sp. 1(-2) celled, shorter and narrower 78
78a Sp. 15-24(29) x 2,5-3,2 µm, oil 3(+), hymenium yellow(ish), drying orange-ochraceous, Apo.
± pulvinat, up to 6 mm , thick, KOH-reaktion yellow (always? not in exs.?) M. retincola

78b Sp. 8-12(15) x 2,2-3,2 µm, oil 1-2, hymenium grey, blackish-grey, apo.  flat, patelliform,


KOH-Reaktion only in fresh collections short but distinctly yellow, in overmature or dried
collections normally not longer visible, marginal cells vesiculare, inconspicuous

M. hydrophila

also collections without any trace of subicular hyphae and x or cristals and x or slightly smaller and narrower spores (transition vs. M. palustris)

identical is M. epithypha Karsten.
80a Sp. (4)4,5-6(7) x 0,8-1,2 µm, oil 0-0,5, in KOH some paraphyses orange-yellow
discolorating (= KOH +?), paraphyses narrower than the asci (difference to M. caricina!)

M. phragmitis

80b Sp. 6,5-8,5 x (1)1,2-1,5 µm, oil 0,5-1, consisting of 1 small and a few very tiny droplets,


scattered through the whole spore (exsiccate, living too?), marginal cells conspicuous,
brownish up to the margin, end cells claviform, 15-22 x 5-8 µm M. evilescens (TYPE)

Identical is M. simillima Karsten.

80c Sp. 6,5-8,5 x 1,8-2,2 µm, oil 0-1 (few tiny droplets), marginal cells subhyaline,
vesiculare, up to ca. 10 x 8 µm (growth on Phragmites to verify) M. palustris (TYPE)

The problems within this complex are big: on the one hand the types are quite distinct, on the other hand are recent findings of these „species“ not too rare and the hiatus between the types disappeares. Until now I have seen only one or two recent collection which corresponds to the type of M. evilescens appr. 100%, but many collections which have to be located between M. evilescens and M. palustris.



Le Gal & Mangenot describe a (lecto-)type of M. palustris, which probably belongs better to M. evilescens (small spores, marginal cells)

May be not distinct on species level!
85a Sp. (4)4,5-6(7) x 0,8-1,2 µm, oil 0-0,5, in KOH some paraphyses orange-yellow
discolorating (= KOH +?), paraphyses 2,5-3 µm wide, narrower than the asci

M. phragmitis

85b Sp. 4-5,5 x 1 µm, paraphyses 3,5 µm wide, slightly broader than the asci, KOH +



M. caricina

85c Paraphyses narrower than the asci or sp. otherwise 86


86a Sp. multiseptate within the living asci 88

86b Sp. not septate, except sometimes before germinating and then not in living asci 100


88a Ascus reaction rr 90

88b Ascus reaction bb 94


90a In mature, turgescent asci sp. 4-6-celled, 30-42 x 4,3-5,2 µm M. (“Niptera”) pulla

Exs. Jaklitsch: sp. 27-34 x 4,5-5,2 µm

90b In mature, turgescent asci sp. 2-4-celled, < 4,2 µm broad 92
92a Sp. (25)30-40 x 3-4 µm, (2)4-celled, constricted at the septae, asci up to 160 µm long,
ect. exc. dark brown M. (“Niptera”) pilosa

92b Sp. 36-46 x 2,5-3,5 µm, 4-celled, not constricted at the septae, asci up to 80-90 µm long,


ect. exc. ochraceous (acccording to Baral) "Niptera" filispora
94a Sp. 35-60 x 2,5-4 µm 95

94b Sp. smaller 96


95a Asci without croziers, sp. 47-53 x 3,5-4 µm, oil 4,5-5 "Belonium" asteroma

95b Asci with croziers, Sp. 35-55(60) x 2,2-3 µm, oil 1-2 M. iridis

Acc. to Baral (in vivo veritas 2005) = Trichobelonium guestphalicum.

Has to be renamed in Mollisia: M. iridis (Crouan & Crouan 1867) LeGal 1953 is inval. because of M. iridis (Rehm 1882) Saccardo 1889


96a KOH-reaction yellow, sp. in IKI with red reaction (gelified) sheath "Niptera" lacustris

96b KOH-reaction negativ, sp. without red reaction in IKI 97


97a Sp. 16-17 x 2,5-3 µm or 19-23 x 3 µm, Öl 3 cf. M. luctuosa

97b Sp. 18-22 x 5-7,5 µm, länglich-oval, schon im Ascus mit 1 dicken Septum, Öl 2,


Paraphysen fädig und etwas gabelig-verzweigt "Nimbomollisia" eriophori
100a IKI negativ, medulla with cristals, KOH strong yellow, sp. 14-17 x 2,8-3 µm (another
collection 19-24 x 2,5-3,2 µm) M. phalaridis

100b Ascus IKI rr 101

100c Ascus IKI bb 102
101a Sp. 12-16 x 2-2,2 µm (exs.), oil 2-3, on ?Juncus in peat bog M. "spec.006"

101b Sp. 8-13,5(16) x 1,8-2,5 µm bzw. (9)10-13,5(15) x 1,8-2,3 µm (exs.), oil somewhat less?,


leave bases of Sesleria varia in Teucrio-Seslerietum M. lothariana

Both taxa are obviousely very similar if not identical (hymenium dry crème- to orange-yellow, anchoring hyphae only 1,5-2,5 µm breit, etc.), only the ecology differs strongly.


102a Medulla full of cristals, KOH reaction (short time) strong (only fresh ?!), sp. 8-12(15) x
2,2-3,2 µm, oil 0,5-1(2) M. hydrophila

Identical is M. epithypha Karsten.

SV NH4OH yellow, sp. 6,5-10 x 1,5-1,8 µm [dead], slightly scutuloid, oil 0

M. citrinopigmentosa

SV Sp. 6-7,5 x 1,5-2 [dead], oil 0, with amyloid granula in the hymenium (probably KOH -)



M. amyloidea

SV Sp. 6,5-10,5 x 2 [living], oil 0 (probably KOH -) M. epityphicola

102b No cristals, KOH-reaction negative or spores smaller and narrower 103
103a Sp. 12-20 x 2,5-3,5 µm M. luctuosa

103b Sp. smaller 105


105a Sp. 4-8 x 1-1,3, KOH-reaction yellow, paraphyses as wide as the asci M. caricina

105b Sp. wider, KOH-reaction negativ (all following species?), paraphyses more narrow than


the asci 106
106a Sp. 7-10(11) x 1,5-2,2 µm, KOH - M. palustris

106b Sp. 8-12(13) x (2,2)2,5-2,8 µm, one end more pointed than the other, oil 0,5(-1),


subicular hyphae strongly swelling in KOH up to 7-8 µm “Tapesia cinerea

106c Sp. 8,5-11 x 2-2,5 µm, marginal cells multicellular, up to appr. 40 µm lang, end cell


utriform to cylindrical “Tapesia eriophori

106d Sp. 9,5-11 x 2-2,8 µm, KOH -, oil 1-2, ect. Exc. a dark textura prismatica



M. juncina (ss. Br & Kr)

most probably no Mollisia species

SV Sp. 6,5-10,5 x 2 [living], slightly scutuloid, oil 0 M. epityphicola

SV Sp. 5-12 x 2-2,5 µm [dead], oil 0, with a covering of amyloid granula over the hymenium



M. variabilispora

SV Sp. 6-7,5 x 1,5-2 [dead], oil 0, with amyloid granula in the upper part of the hymenium



M. amyloidea

LEAFES
120a Sp. 10-12 x 3-3,5 µm, marginal cells club-shaped, not agglutinated, on Rubus (only ?)

M. gabretae

120b Sp. 9-12 x -2,5 µm, mature often septate, marginal cells nearly cylindrical, sometimes
agglutinated (is this a character?), on Quercus (above all Q. ilex) in (sub-)mediterranean
to atlantic areas (only?) M. muelleri-argovensis

120c Sp. smaller 121


121a Sp. 5,5-8 x 1,7-2,2 µm, marginal cells 2-celled zweizellig, roundish to pear-shaped, not
agglutinated M. rabenhorstii

121b Sp. 5-7 x 1,2-1,5(1,8) µm, marginal cells small, conspicuous, brownish, agglutinated to


triangular teeth M. lithocarpi

MOSSES
140 ---

SV Sp. 13-16 x 2,5-3,5 µm, septate (in the ascus?), KOH-reaction unknown, marginal cells


subglobos and with brownish incrustations, ascus reaction bb, on living mosses

M. polytrichicola

========================================================================




TYPUS-UNTERSUCHUNGEN
Normaldruck nicht zu Mollisia gehörend.

Fettdruck Art gehört zu Mollisia s. l., wird aber als synonym angesehen.

Fettdruck, unterstrichen Art gehört zu Mollisia, Epithet wird als anwendbar betrachtet.
Crustula corylacea Velenovsky: Holotypus PRM 150537, PRM (= M. ligni)

Crustula nigra Velenovsky: Holotypus PRM 614736, PRM (= Durella nigra comb. nov. ined.)

Crustula quercina Velenovsky: Lektotypus PRM 153167, PRM (= M. ligni)
M. angelicae Dearness: Sydow 744 (Fungi exsiccati exotici), HUH

M. anserina Velenoský: Holotypus PRM 152223, PRM (= M. palustris)

M. arescens Rehm: Ule 1184, HUH

M. asclepiadis Ellis & Everh.: Can. Fungi, HUH

M. betulina Velenoský: Holotypus PRM 152300, PRM (= cf. Pyrenopeziza aquosa)

M. caespiticia Karsten: Lectotypus PAK 3588, H, Syntypus PAK 3589, H

M. dakotensis Rehm: Authentic Specimen N. Dakota Fungi, HUH

M. ephemera Rehm: Ule 1277, HUH

M. epithypa Karsten: PAK 4396, H ; als „epitypha n. sp.“, könnte Typus sein.

M. evilescens Karsten: Neotypus PAK 3592, H

M. faginea Velenovsky: Holotypus PRM 151676, PRM

M. gallincola Velenovsky: Lektotypus PRM 148261, PRM (unklar, vielleicht eigenständig)

M. hydrophila Karsten: Holotypus Fung. Fenn. Exs. Nr. 643, H

M. laeta Rick: Typus + Syntypus (nicht separat gekennzeichnet), HUH

M. lilacina Clement: authentic specimen ex Herb. Shear, HUH

M. lithocarpi Cash: Isotypus, UPLB 9109 , BR .....

M. lothariana Gminder: Holotypus 137/1997 herb. Krieglsteiner et filii, STU

M. mikaniae Rehm: Ule 918, HUH

M. papillata Earle: Plants of Pac. Coast No. 207, HUH

M. perparvula Karsten: Lectotypus PAK 2922, H

M. petiolorum Cash: Hawaiian Fungi No. 555, HUH

M. phaea Rehm: Asc. 713, HUH

M. pilifera Velenoský: Holotypus PRM 152217, PRM (= Hyaloscypha spec.)

M. revincta Karsten: Typus PAK 3605, H

M. rhinanthi Karsten: Typus PAK 3652 + 3653, H

M. simillima Karsten: Holotypus PAK 4530, H

M. spraguei Santesson: Holotypus, HUH
Pseudoniptera quercina Velenovsky: Lektotypus PRM 152904, PRM (= Hymenoscyphus spec.)
Tapesia airae Velenovsky: Holotypus PRM 154076, PRM (= M. cf. phragmitis)

Tapesia alpina Velenovsky: Lektotypus PRM 148518, PRM (M. velenovskyi nom. nov.)

Tapesia cinerea Velenovsky: Lektotypus PRM 153121, PRM (M. palustris complex)

Tapesia dimorpha Velenovsky: Holotypus PRM 153177, PRM (M. cf. olivaceocinerea)

Tapesia eriophori Velenovsky: Lektotypus PRM 147734, PRM (M. palustris complex)

Tapesia exigua Velenovsky: Holotypus PRM 153031, PRM (empty)

Tapesia globulifera Velenovsky: Holotypus PRM 153169, PRM

Tapesia ladae Velenovsky: Holotypus PRM 1531728, PRM

Tapesia lentiformis Velenovsky: Holotypus PRM 153104, PRM (???)

Tapesia ochroleuca Velenovsky: Holotypus PRM 148710, PRM

Tapesia peruni Velenovsky: Lektotypus PRM 812443, PRM

Tapesia pezizellaeformis Velenovsky: Holotypus PRM 153038, PRM (= Hyaloscypha daedalea)

Tapesia phragmitis Velenovsky: Lektotypus PRM 147397, PRM

Tapesia sesleriae Velenovsky: Holotypus PRM 148511, PRM (nomen dubium oder Cistella)

Tapesia urnigera Velenovsky: Holotypus PRM 153104, PRM (???,M. cf. phragmitis)
Trichobelonium caricinum Velenovský: Holotypus PRM 148431, PRM (= M. pilosa/filispora)

BR National Botanic Garden of Belgium, Meise

H Herbarium of the University of Helsinki.

HUH Herbarium of the Harvard-University in New York

M Botanische Staatssammlung München

MANCH Manchester Museum, University of Manchester

PAK Herbarium of Petter Adolf Karsten (in H)

PRM Prague, Herbarium of Velenovský



UPLB University of the Philippines at Los Baños College, Laguna

WU Herbarium of the University of Vienna


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