Odonata references




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ODONATA REFERENCES
These references are associated with the books Dragonflies and Damselflies of the West, 2009, and Dragonflies and Damselflies of the East, 2011, by Dennis Paulson, Princeton University Press.
The references will be primarily to publications on the natural history of the species, but some recent taxonomic papers will be included, mostly at the generic level. The references will furnish much more information than could be included in the book. References to larval ecology are not included. Information about many species can also be found in the general works cited in the List of References in the books themselves.
References added through 2007; updates planned.
JEWELWINGS Calopteryx

Reference Johnson 1974.
Sparkling Jewelwing Calopteryx dimidiata

References Waage 1984, 1988a.
River Jewelwing Calopteryx aequabilis

References Conrad & Herman 1987, Henderson & Herman 1984.
Ebony Jewelwing Calopteryx maculata

References Alcock 1979, 1983, Ballou 1984, Erickson & Reid 1989, Forsyth & Montgomerie 1987, Johnson 1962c, Waage 1972, 1973, 1978, 1979, 1980, 1984, 1987, 1988b.
RUBYSPOTS Hetaerina

References Garrison 1990, Johnson 1973.
American Rubyspot Hetaerina americana

References Bick & Sulzbach 1966, Grether 1996a, 1996b, Grether & Grey 1996, Grether & Switzer 2000, Johnson 1961, 1962a, 1963, McCafferty 1979.
Canyon Rubyspot Hetaerina vulnerata

Reference Alcock 1982.
Smoky Rubyspot Hetaerina titia

References Harp 1986, Johnson 1961, 1963.
Great Spreadwing Archilestes grandis

Reference Bick & Bick 1970.
California Spreadwing Archilestes californicus

Reference Kennedy 1915.
Chalky Spreadwing Lestes sigma

Reference Johnson 1975.
Plateau Spreadwing Lestes alacer

References Bird 1933, Larsen 2007.
Spotted Spreadwing Lestes congener

References Cannings et al 1980, McMillan & Arnold 2004, Sawchyn & Gillott 1974a.
Northern Spreadwing Lestes disjunctus

References Donnelly 2003, Sawchyn & Gillott 1974b, Walker 1952.
Southern Spreadwing Lestes australis

References Bick & Bick 1961, Donnelly 2003, Walker 1952.
Sweetflag Spreadwing Lestes forcipatus

References Donnelly 2003, Walker 1952.
Lyre-tipped Spreadwing Lestes unguiculatus

References Bick & Hornuff 1965, Sawchyn & Gillott 1974b.
Slender Spreadwing Lestes rectangularis

Reference Gower & Kormondy 1963.
Emerald Spreadwing Lestes dryas

Reference Sawchyn & Gillott 1974b.
Prairie Bluet Coenagrion angulatum

Reference Sawchyn & Gillott 1975.
Taiga Bluet Coenagrion resolutum

Reference Sawchyn & Gillott 1975.
AMERICAN BLUETS Enallagma

References Garrison 1984, May 2002.
Arroyo Bluet Enallagma praevarum

References Johnson 1964b, 1964c.
River Bluet Enallagma anna

References Kennedy 1917, La Rivers 1940.
Claw-tipped Bluet Enallagma semicirculare

Reference Garrison & May 1988.
Familiar Bluet Enallagma civile

References Bick & Bick 1963, Bird 1933, Johnson 1964b, Moss 1992.
Azure Bluet Enallagma aspersum

Reference Bick & Hornuff 1966.
Alkali Bluet Enallagma clausum

Reference Kennedy 1917.
Northern Bluet Enallagma annexum

References Doerksen 1980, Garrison 1978.
Boreal Bluet Enallagma boreale

Reference Anholt 1992.
Hagen’s Bluet Enallagma hageni

References Fincke 1982, 1986.
Baja Bluet Enallagma eiseni

Reference Manolis et al 2006.
Stream Bluet Enallagma exsulans

Reference Bick & Hornuff 1966.
Slender Bluet Enallagma traviatum

References Donnelly 1973, Robinson 1981.
Orange Bluet Enallagma signatum

Reference Bird 1933.
FORKTAILS Ischnura

References Chippendale et al 1998, Donnelly 1965, Robinson & Allgeyer 1996, Robinson & Novak 1997.
Swift Forktail Ischnura erratica

References Cannings & Doerksen 1979, Paulson & Cannings 1980.
Lilypad Forktail Ischnura kellicotti

Reference Robinson & Jordan 1996.
Rambur’s Forktail Ischnura ramburii

Reference Robertson 1985.
Pacific Forktail Ischnura cervula

References Dickerson et al 1992, Kennedy 1915.
Plains Forktail Ischnura damula

Reference Johnson 1964a.
Mexican Forktail Ischnura demorsa

Reference Johnson 1966
Western Forktail Ischnura perparva

Reference Kennedy 1915.
Eastern Forktail Ischnura verticalis

References Fincke 1987, Grieve 1937.
Black-fronted Forktail Ischnura denticollis

References Córdoba-Aguilar 1992, 1993, Kennedy 1917, Leong and Hafernik 1992.
San Francisco Forktail Ischnura gemina

References Garrison & Hafernik 1981a, 1981b, Hafernik & Garrison 1986, Kennedy 1917, Leong & Hafernik 1992.
Fragile Forktail Ischnura posita

References Novelo-Gutiérrez & Peña-Olmedo 1989, Robinson et al 1985.
Painted Damsel Hesperagrion heterodoxum

Reference De Marmels 2002.
Western Red Damsel Amphiagrion abbreviatum

Reference Pritchard and Kortello 1997.
Aurora Damsel Chromagrion conditum

References Bick, Bick, & Hornuff 1976, De Marmels 2002.
Exclamation Damsel Zoniagrion exclamationis

References De Marmels 2002, Kennedy 1917.
Desert Firetail Telebasis salva

Reference Robinson & Frye 1986.
Duckweed Firetail Telebasis byersi

Reference Lounibos et al 1990.
Sedge Sprite Nehalennia irene

References Forbes et al 1995, Hilton 1983c.
DANCERS Argia

Reference Garrison 1994b.
Blue-fronted Dancer Argia apicalis

References Bick & Bick 1965a, 1965b.
Powdered Dancer Argia moesta

References Bick & Bick 1972, Borror 1934.
Blue-ringed Dancer Argia sedula

Reference Robinson et al 1983.
Variable Dancer Argia fumipennis

Reference Gloyd 1968.
Aztec Dancer Argia nahuana

Reference Bick & Bick 1958.
Yaqui Dancer Argia carlcooki

Reference Behrstock et al 2004.
Pima Dancer Argia pima

Reference Hoekstra & Smith 2000.
Sabino Dancer Argia sabino

Reference Hoekstra & Smith 2000.
Springwater Dancer Argia plana

References Bick & Bick 1958, 1968, 1971, 1972.
Vivid Dancer Argia vivida

References Conrad & Pritchard 1988, 1989, 1990, Garrison 1978, Kennedy 1915, Pritchard and Kortello 1997.
Emma's Dancer Argia emma

Reference Kennedy 1915.
SHADOWDAMSELS Palaemnema

Reference Calvert 1931.
Desert Shadowdamsel Palaemnema domina

Reference Hoekstra & Garrison 1999.
Orange-striped Threadtail Protoneura cara

Reference González-Soriano 2001.
Gray Petaltail Tachopteryx thoreyi

References Barlow 1991, Dunkle 1981, Walker & Smentowski 2003.
Black Petaltail Tanypteryx hageni

References Clement & Meyer 1980, Meyer & Clement 1978, Svihla 1959.
Harlequin Darner Gomphaeschna furcillata

Reference Kennedy 1936.
Fawn Darner Boyeria vinosa

Reference Paulson 1999.
Cyrano Darner Nasiaeschna pentacantha

Reference Paulson 1999.
Riffle Darner Oplonaeschna armata

Reference Johnson 1968.
TWO-SPINED DARNERS Gynacantha

Reference Williamson 1923a.
Twilight Darner Gynacantha nervosa

References Paulson 1999, Williams 1937.
Bar-sided Darner Gynacantha mexicana

Reference Paulson 2006.
THREE-SPINED DARNERS Triacanthagyna

References Williamson 1923a, von Ellenrieder and Garrison 2003.
Regal Darner Coryphaeschna ingens

Reference Paulson 1999.
MOSAIC DARNERS Aeshna

Reference Walker 1912.
Variable Darner Aeshna interrupta

Reference Kennedy 1917.
Zigzag Darner Aeshna sitchensis

Reference Cannings 1982.
Walker’s Darner Aeshna walkeri

Reference Kennedy 1917.
Persephone’s Darner Aeshna persephone

Reference Donnelly 1961.
NEOTROPICAL DARNERS Rhionaeschna

Reference von Ellenrieder 2003.
Blue-eyed Darner Rhionaeschna multicolor

Reference La Rivers 1938.
Blue-spotted Comet Darner Anax concolor

References Geijskes 1968, Paulson 1966.
Comet Darner Anax longipes

Reference Paulson 1999.
Giant Darner Anax walsinghami

Reference Córdoba-Aguilar 1995.
Common Green Darner Anax junius

References Freeland et al 2003, Kennedy 1917, Paulson 1999, Trottier 1966, Young 1967.
CLUBTAIL FAMILY GOMPHIDAE

Reference Carle 1986.
Gray Sanddragon Progomphus borealis

Reference Kennedy 1917.
Common Sanddragon Progomphus obscurus

Reference Reinhardt 2003.
GREATER FORCEPTAILS Aphylla

Reference Garrison 1986.
Two-striped Forceptail Aphylla williamsoni

References Gloyd 1936, Paulson 1999.
COMMON CLUBTAILS Gomphus

Reference May & Carle 1996.
Pacific Clubtail Gomphus kurilis

Reference Kennedy 1917.
Pronghorn Clubtail Gomphus graslinellus

Reference Reinhardt 2003.
Subgenus Hylogomphus

Reference Carle 1979.
Ozark Clubtail Gomphus ozarkensis

Reference Susanke & Harp 1991.
Plains Clubtail Gomphus externus

Reference Reinhardt 2003.
Tamaulipan Clubtail Gomphus gonzalezi

Reference Dunkle 1992.
Columbia Clubtail Gomphus lynnae

Reference Paulson 1983.
Olive Clubtail Stylurus olivaceus

Reference Kennedy 1917.
Russet-tipped Clubtail Stylurus plagiatus

Reference Paulson 1999.
Brimstone Clubtail Stylurus intricatus

Reference Kennedy 1917.
RINGTAILS Erpetogomphus

Reference Garrison 1994a.
SNAKETAILS Ophiogomphus

Reference Carle 1992, Kennedy 1917.
Great Basin Snaketail Ophiogomphus morrisoni

References La Rivers 1946, Paulson 1998b.
Interior Least Clubtail Stylogomphus sigmastylus

Reference Cook & Laudermilk 2004.
Grappletail Octogomphus specularis

Reference Kennedy 1917.
SPIKETAILS Cordulegaster

Reference Carle 1983.
Pacific Spiketail Cordulegaster dorsalis

References Cruden 1969, Kennedy 1917.
Apache Spiketail Cordulegaster diadema

Reference Alcock 1985.
RIVER CRUISERS Macromia

Reference Williamson 1909.
Swift River Cruiser Macromia illinoiensis

Reference Donnelly & Tennessen 1994.
Gilded River Cruiser Macromia pacifica

Reference Williams 1979.
Royal River Cruiser Macromia taeniolata

Reference Paulson 1999.
Western River Cruiser Macromia magnifica

References Kennedy 1915, Walker 1937.
Ebony Boghaunter Williamsonia fletcheri

Reference Charlton & Cannings 1993.
American Emerald Cordulia shurtleffii

Reference Hilton 1983a.
STRIPED EMERALDS Somatochlora

Reference Walker 1925.
Treeline Emerald Somatochlora sahlbergi

Reference Cannings & Cannings 1985.
Mountain Emerald Somatochlora semicircularis

Reference Kennedy 1913.
Plains Emerald Somatochlora ensigera

Reference Williamson 1922b.
Mocha Emerald Somatochlora linearis

Reference Williamson 1922b.
Texas Emerald Somatochlora margarita

Reference Donnelly 1962.
Clamp-tipped Emerald Somatochlora tenebrosa

Reference Williamson 1922b.
Orange Shadowdragon Neurocordulia xanthosoma

Reference Williams 1976.
BASKETTAILS Epitheca

References Davis 1933, May 1995, Walker 1966.
Beaverpond Baskettail Epitheca canis

Reference Kennedy 1917.
Prince Baskettail Epitheca princeps

Reference Paulson 1999.
Common Whitetail Plathemis lydia

References Campanella & Wolf 1974, Jacobs 1955, Koenig 1990, 1991, Koenig & Albano 1985, 1987, McVey 1984.
CORPORALS Ladona

References Bennefield 1965, May 1992.
Blue Corporal Ladona deplanata

Reference Paulson 1999.
Chalk-fronted Corporal Ladona julia

Reference Hilton 1983b.
KING SKIMMERS Libellula

References Artiss et al 2001, Carle & Kjer 2002, Kambhampati et al 1999, Kennedy 1922.
Four-spotted Skimmer Libellula quadrimaculata

Reference Convey 1989.
Neon Skimmer Libellula croceipennis

Reference Williams 1977.
Flame Skimmer Libellula saturata

References Alcock 1989a, DeBano 1993, 1996, La Rivers 1938.
Bleached Skimmer Libellula composita

Reference Beckemeyer 2004.
Hoary Skimmer Libellula nodisticta

Reference Kennedy 1917.
Twelve-spotted Skimmer Libellula pulchella

Reference Pezzalla 1979.
Widow Skimmer Libellula luctuosa

References Campanella 1975, Garrison 1976, Moore 1987a, 1987b, 1990.
Bar-winged Skimmer Libellula axilena

Reference Dunkle 1985.
Slaty Skimmer Libellula incesta

References Dunkle 1985, Paulson 1999.
Great Blue Skimmer Libellula vibrans

References Dunkle 1985, Paulson 1999.
Golden-winged Skimmer Libellula auripennis

Reference Paulson 1999.
Needham’s Skimmer Libellula needhami

Reference Paulson 1999.
Carmine Skimmer Orthemis discolor

References Novelo and González 1984, Paulson 1998a, Young 1980.
Roseate Skimmer Orthemis ferruginea

References Novelo and González 1984 (may refer mostly to Carmine), Paulson 1999.
AMBERWINGS Perithemis

Reference Ris 1930.
Eastern Amberwing Perithemis tenera

References Hardy 1966, Jacobs 1955, Paulson 1999, Schultz & Switzer 2001, Schiffer 1968, Switzer 1997, 2002a, 2002b, Switzer & Walters 1999.
Red-tailed Pennant Brachymesia furcata

Reference Paulson 1999.
Four-spotted Pennant Brachymesia gravida

Reference Paulson 1999.
SMALL PENNANTS Celithemis

Reference Williamson 1922a.
Ornate Pennant Celithemis ornata

Reference Paulson 1999.
Calico Pennant Celithemis elisa

Reference Waage 1986.
Banded Pennant Celithemis fasciata

Reference Paulson 1999.
Halloween Pennant Celithemis eponina

References Miller 1982, Paulson 1999.
WHITEFACES Leucorrhinia

References Hovmöller & Johansson 2004, Westmann et al 2000.
Dot-tailed Whiteface Leucorrhinia intacta

References Wolf & Waltz 1984, Wolf et al 1997.
PONDHAWKS Erythemis

References Kennedy 1923, Williamson 1923b.
Pin-tailed Pondhawk Erythemis plebeja

Reference De Marco et al 2002.
Great Pondhawk Erythemis vesiculosa

Reference Paulson 1999.
Eastern Pondhawk Erythemis simplicicollis

References Currie 1963, Donnelly 2004, Lounibos et al 1990, May & Baird 2002, McVey 1985, McVey and Smittle 1984, Paulson 1999, Waage 1986.
Western Pondhawk Erythemis collocata

Reference Donnelly 2004.
DRAGONLETS Erythrodiplax

Reference Borror 1942.
Red-faced Dragonlet Erythrodiplax fusca

Reference Paulson 2003.
Plateau Dragonlet Erythrodiplax basifusca

Reference Paulson 2003.
Little Blue Dragonlet Erythrodiplax minuscula

References Paulson 1999, 2003.
Seaside Dragonlet Erythrodiplax berenice

Reference Paulson 1999.
Band-winged Dragonlet Erythrodiplax umbrata

Reference Paulson 1999.
Filigree Skimmer Pseudoleon superbus

Reference Córdoba-Aguilar 2003.
MEADOWHAWKS Sympetrum

References Carle 1993, Pilgrim & von Dohlen 2007.
Variegated Meadowhawk Sympetrum corruptum

References Arnaud 1972, La Rivers 1938.
Cardinal Meadowhawk Sympetrum illotum

Reference Kennedy 1917.
Red-veined Meadowhawk Sympetrum madidum

Reference Cannings 1980.
Cherry-faced Meadowhawk Sympetrum internum

Reference Schmidt 1987.
Band-winged Meadowhawk Sympetrum semicinctum

References Pilgrim & von Dohlen 2007, Walker 1951.
Black Meadowhawk Sympetrum danae

References Michiels & Dhondt 1989, 1990.
Autumn Meadowhawk Sympetrum vicinum

References May 1998, McMillan 1996.
Spot-winged Meadowhawk Sympetrum signiferum

References Cannings & Garrison 1991, Pilgrim & von Dohlen 2007, Upson 2000.
TROPICAL DASHERS Micrathyria

Reference Paulson 1969.
Spot-tailed Dasher Micrathyria aequalis

References May 1977, 1980.
Blue Dasher Pachydiplax longipennis

References Baird & May 1997, 2003, Johnson 1962b, McKinnon & May 1994, Paulson 1999, Penn 1951, Robey 1975, Sherman 1983.
Red Rock Skimmer Paltothemis lineatipes

References Alcock 1987, 1989b, 1990, Dunkle 1978.
Jade-striped Sylph Macrothemis inequiunguis

Reference Gallucci 2007.
Masked Clubskimmer Brechmorhoga pertinax

Reference Alcock 1989b.
Evening Skimmer Tholymis citrina

Reference Miller & Miller 1985 (related species).
Hyacinth Glider Miathyria marcella

Reference Paulson 1999.
Antillean Saddlebags Tramea insularis

Reference Paulson 1999.
Red Saddlebags Tramea onusta

Reference Young 1967.
Carolina Saddlebags Tramea carolina

Reference Paulson 1999.
Black Saddlebags Tramea lacerata

Reference Kennedy 1917.
Wandering Glider Pantala flavescens

Reference Paulson 1999.
Spot-winged Glider Pantala hymenaea

References La Rivers 1938, Paulson 1999.
Marl Pennant Macrodiplax balteata

Reference Paulson 1999.
List of References
Alcock, J. 1979. Multiple mating in Calopteryx maculata (Odonata: Calopterygidae) and the advantage of non-contact guarding by males. Journal of Natural History 13: 439-446.

Alcock, J. 1982. Postcopulatory mate guarding by males of the damselfly Hetaerina vulnerata Selys (Odonata: Calopterygidae). Animal Behaviour 30: 99-107.

Alcock, J. 1983. Mate guarding and the acquisition of new mates in Calopteryx maculata (P. de Beauvois) (Zygoptera: Calopterygidae). Odonatologica 12: 153-159.

Alcock, J. 1985. Reproductive behavior of Cordulegaster diadema Selys (Anisoptera: Cordulegastridae). Odonatologica 14: 313-317.

Alcock, J. 1987. Male reproductive tactics in the libellulid dragonfly Paltothemis lineatipes: temporal partitioning of territories. Behavior 103: 157-173.

Alcock, J. 1989a. The mating system of Libellula saturata Uhler (Anisoptera: Libellulidae). Odonatologica 18: 89-93.

Alcock, J. 1989b. Annual variation in the mating system of the dragonfly Paltothemis lineatipes (Anisoptera: Libellulidae). Journal of the Zoological Society of London 218: 597-602.

Alcock, J. 1989b. The mating system of Brechmorhoga pertinax (Hagen): the evolution of brief patrolling bouts in a “territorial” dragonfly (Odonata: Libellulidae). Journal of Insect Behavior 2: 49-62.

Alcock, J. 1990. Oviposition resources, territoriality and male reproductive tactics in the dragonfly Paltothemis lineatipes (Odonata: Libellulidae). Behaviour 113: 251-263.

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