M. A. Institute of Archaeology, Tel Aviv University. Including complementary studies for a B. A. awarded a

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Contact:  levyadun@research.haifa.ac.il

Born November 19, 1952

Place of Birth Israel.

Military Service 1971-1974 (disabled veteran, lost a leg in the 1973 war).

1994-1997, Scientist, Rank Major

  1. B.Sc., Biology, Faculty of Life Sciences, Tel Aviv University.

  2. M.Sc., Department of Botany, Tel Aviv University (awarded a summa cum laude).

  3. M.A. Institute of Archaeology, Tel Aviv University. Including complementary studies for a B.A. (awarded a

  4. summa cum laude).

  5. Ph.D., Department of Botany, Tel Aviv University.

  6. Postdoctoral fellow, Department of Plant Genetics, The Weizmann Institute of Science.

Major Research Interests

  1. Biology and ecology of trees and other woody plants.

  2. Developmental processes in plants: meristematic activity and its ecology and regulation, fate of apical meristems, homeosis, pattern formation.

  3. Palaeoecology: beginning of agriculture, dendrochronology, identification of botanical material from archaeological excavations, reconstruction of palaeoenvironments, the influence of human activity on the ecosystem.

  4. The ecology and evolution of colors in plants.

  5. Arabidopsis thaliana as a model for wood and fiber formation.


  1. Plant physiology. 

  2. Introductionary and advanced field courses in geobotany and plant ecology.

  3. Plants and Human Culture.

  4. Plants structure and function.


Refereed Articles in Journals

Biology Journals

  1. Liphschitz, N., S. Lev-Yadun & Y. Waisel. 1981.The annual rhythm of activity of the lateral meristems (cambium and phellogen) in Cupressus sempervirens L.Annals of Botany 47:485-496.

  2. Liphschitz, N., S. Lev-Yadun, E. Rosen & Y. Waisel. 1984.The annual rhythm of activity of the lateral meristems (cambium and phellogen). in Pinus halepensis Mill. and Pinus pinea L.IAWA Bulletin n.s. 5:263-274.

  3. Liphschitz, N., S. Lev-Yadun & Y. Waisel. 1985.The annual rhythm of the lateral meristems (cambium and phellogen) in Pistacia lentiscus L.IAWA Bulletin n.s. 6:239-244.

  4. Liphschitz, N. & S. Lev-Yadun. 1986.Cambial activity of evergreens and seasonal dimorphics around the Mediterranean.IAWA Bulletin n.s. 7:145-153.

  5. Lev-Yadun, S. & N. Liphschitz. 1987.The ontogeny of gender in Cupressus sempervirens L.Botanical Gazette 148:407-412.

  6. Lev-Yadun, S. & N. Liphschitz. 1989.Sites of first phellogen initiation in conifers.IAWA Bulletin n.s. 10:43-52.

  7. Lev-Yadun, S. & R. Aloni. 1990.Vascular differentiation in branch junctions of trees: circular patterns and functional significance. Trees Structure and Function 4:49-54.

  8. Lev-Yadun, S. & R. Aloni. 1990.Polar patterns of periderm ontogeny, their relationship to leaves and buds, and the control of cork formation.IAWA Bulletin n.s. 11:289-300.

  9. Lev-Yadun, S. & R. Aloni. 1991.Polycentric vascular rays in Suaeda monoica and the control of ray initiation and spacing.Trees Structure and Function 5:22-29.

  10. Lev-Yadun, S. & R. Aloni. 1991.Wound-induced periderm tubes in the bark of Melia azedarach, Ficus sycomorus and Platanus acerifolia. IAWA Bulletin n.s. 12:62-66.

  11. Lev-Yadun, S. & R. Aloni. 1991.Natural and experimentally induced dispersion of aggregate rays in shoots of Quercus ithaburensis Decne. and Q. calliprinos Webb.Annals of Botany 68:85-91.

  12. Lev-Yadun, S. & R. Aloni. 1991.An experimental method of inducing "hazel" wood in Pinus halepensis (Pinaceae).IAWA Bulletin n.s. 12:445-451.

  13. Lev-Yadun, S. 1992.Abnormal cones in Cupressus sempervirens.Aliso 13:391-394.

  14. Lev-Yadun, S. 1992.Aggregated cones in Pinus halepensis.Aliso 13:475-485.

  15. Lev-Yadun, S. & R. Aloni. 1992.Experimental induction of dilatation meristem in Melia azedarach.Annals of Botany 70:379-386.

  16. Lev-Yadun, S. & R. Aloni. 1992.The role of wounding in the differentiation of vascular rays.International Journal of Plant Sciences 153:348-357.

  17. Lev-Yadun, S. & R. Aloni. 1993.Bark structure and mode of canopy regeneration in trees of Melia azedarach L.Trees Structure and Function 7:144-147.

  18. Lev-Yadun, S. & R. Aloni. 1993.Effect of wounding on the relations between vascular rays and vessels in Melia azedarach L.New Phytologist 124:339-344.

  19. Lev-Yadun, S. & R. Aloni. 1993.Variant secondary growth in old stems of Ephedra campylopoda C. A. Mey.Botanical Journal of the Linnean Society 112:51-58.

  20. Eyal, Y., Y. Meller, S. Lev-Yadun & R. Fluhr. 1993.A basic- type PR-1 promoter directs ethylene responsiveness, vascular and abscission zone-specific expression.Plant Journal 4:225-234.

  21. Lev-Yadun, S. 1994.Experimental evidence for the autonomy of ray differentiation in Ficus sycomorus L.New Phytologist 126:499-504.

  22. Lev-Yadun, S. 1994.Radial fibres in aggregate rays of Quercus calliprinos Webb. -- evidence for radial signal flow. New Phytologist 128:45-48 (INCLUDING A COVER PICTURE).

  23. Lev-Yadun, S. 1994.Induction of sclereid differentiation in the pith of Arabidopsis thaliana (L.) Heynh.Journal of Experimental Botany 45:1845-1849.

  24. Lev-Yadun, S. 1994.Induction of near-vessellessness in Ephedra campylopoda C. A. Mey.Annals of Botany 75:683-687.

  25. Lev-Yadun, S. & M. Weinstein-Evron. 1994.Late Epipalaeolithic wood remains from el-Wad Cave, Mount Carmel, Israel.New Phytologist 127:391-396.

  26. Lev-Yadun, S. 1995.Living serotinous cones in Cupressus sempervirens. International Journal of Plant Sciences 156:50-54.

  27. Lev-Yadun, S. 1995.Short secondary vessel members in branching regions in roots of Arabidopsis thaliana.Australian Journal of Botany 43:435-438.

  28. Lev-Yadun, S. & R. Aloni. 1995.Differentiation of the ray system in woody plants.Botanical Review 61:45-88.

  29. Lev-Yadun, S. 1996.Circular vessels in the secondary xylem of Arabidopsis thaliana (L.) Heynh. (Brassicaceae).IAWA Journal 17: 31-35.

  30. Lev-Yadun, S. 1996. A developmental variant of indentation in the scales of female cones of Cupressus sempervirens L.Botanical Journal of the Linnean Society 121:263-269.

  31. Lev-Yadun, S. 1996.Patterns of dilatation growth in Ficus pumila and Ficus sycomorus.Aliso 14:171-177.

  32. Lev-Yadun, S., M. Artzy, E. Marcus & R. Stidsing. 1996.Wood remains from Tel Nami, a Middle Bronze IIa and Late Bronze IIb port, local exploitation of trees and Levantine cedar trade.Economic Botany 50: 310-317.

  33. Lev-Yadun, S., A. Beharav & S. Abbo. 1996. Evidence for polygenic control of fiber differentiation in spring wheat and its  relationship with the GA-insensitivity locus Rht1.Australian Journal of Plant Physiology 23:185-189.

  34. Baum, G., S. Lev-Yadun, Y. Fridmann, T. Arazi, H. Katsnelson, M. Zik & H. Fromm. 1996.Calmodulin binding to glutamate decarboxylase is required for regulation of glutamate and GABA metabolism and normal development in plants.EMBO Journal 15:2988-2996.

  35. Fridlender, M., S. Lev-Yadun, I. Baburek, K. Angelis & A. A. Levy. 1996.Cell divisions in cotyledons after germination: localization, time course and utilization for a mutagenesis assay. Planta 199:307-313.

  36. Yakir, D., S. Lev-Yadun & A. Zangvil. 1996.El Niño and tree growth near Jerusalem over the last 20 years.Global Change Biology 2:97-101.

  37. Lev-Yadun, S. 1997.Fibres and fibre-sclereids in wild-type Arabidopsis thaliana.Annals of Botany 80:125-129.

  38. Zhu-Shimoni, J. X., S. Lev-Yadun, B. Matthews & G. Galili. 1997. Expression of an aspartate kinase homoserine dehydrogenase gene is subject to specific spatial and temporal regulation in vegetative tissues, flowers, and developing seeds.Plant Physiology 113:695-706.

  39. Tianbao Yang, S. Lev-Yadun, M. Feldman & H. Fromm. 1998. Developmentally regulated organ-, tissue-, and cell-specific expression of calmodulin genes in common wheat.Plant Molecular Biology 37:109-120.

  40. Lev-Yadun, S. 1998.The relationship between growth-ring width and ray density and ray height in cell number in the earlywood of Pinus halepensis and Pinus pinea.IAWA Journal 19:131-139.

  41. Lev-Yadun, S. 1999.Eccentric deposition of secondary xylem in stems of the climber Ephedra campylopoda (Gnetales).IAWA Journal 20:165-170.

  42. Lev-Yadun, S. 1999.Articulated cork in Calotropis procera (Asclepiadaceae).Aliso 18:161-163.

  43. Lev-Yadun, S. & S. Abbo. 1999.Traditional human utilization of A’kub (Gundelia tournefortii L.; Asteraceae), in Israel and the Palestinian Authority area.Economic Botany 53:217-219.

  44. Lev-Yadun, S., A. Beharav, R. Di-nur & S. Abbo. 1999.Gibberellic acid (GA) increases fibre cell differentiation and secondary cell- wall deposition in spring wheat (Triticum aestivum L.) culms.Plant Growth Regulation 27:161-165.

  45. Lev-Yadun, S. 2000.Why are underground flowering and fruiting in Israel much more common than anywhere else in the world?Current Science 79:289.

  46. Lev-Yadun, S. 2000.Whirled grain in wood and topological defects. Journal of Theoretical Biology 205:511-514.

  47. Lev-Yadun, S., N. Atzmon & A. Perevolotsky. 2000.Grafting to save the forests.Trends in Plant Science 5:94.

  48. Lev-Yadun, S., A. Gopher & S. Abbo. 2000.The cradle of agriculture. Science 288:1602-1603.

  49. Avivi, Y., S. Lev-Yadun, N. Morozova, L. Libs, L. Williams, J. Zhao &  G. Grafi. 2000.Clausa, a tomato mutant with a wide range of phenotypic perturbations, displays a cell type-dependent expression of the homeobox gene Let6/Tkn2.Plant Physiology 124:541-551

  50. Lev-Yadun, S. & R. Sederoff. 2000.Pines as model gymnosperms to study evolution, wood formation and perennial growth.Journal of Plant Growth Regulation 19:290-305.

  51. Lev-Yadun, S. 2001.Arabidopsis thaliana - a new crop?Nature Biotechnology 19:95.

  52. Lev-Yadun, S. 2001.Intrusive growth - the plant analog to dendrite and axon growth in animals.New Phytologist 150:508-512.

  53. Lev-Yadun, S. 2001.Aposematic (warning) coloration associated with thorns in higher plants.Journal of Theoretical Biology 210:385-388.

  54. Lev-Yadun, S. 2001.Wound effects arrest wave phenomena in the secondary xylem of Rhamnus alaternus (Rhamnaceae).IAWA Journal 22: 295-300.

  55. Lev-Yadun, S. & M.A. Flaishman. 2001.The effect of submergence on ontogeny of cambium and secondary xylem and on fiber lignification in inflorescence stems of Arabidopsis.IAWA Journal 22:113-123.

  56. Abbo, S., S. Lev-Yadun & G. Ladizinsky. (2001).Tracing the wild genetic stocks of crop plants.Genome 44:309-310.

II.Archaeology or Geology Journals

  1. Liphschitz, N., S. Lev-Yadun & Y. Waisel. 1981.Dendroarchaeological investigations in Israel: Masada.Israel Exploration Journal 31:230-234.

  2. Gophna, R., N. Liphschitz & S. Lev-Yadun. 1986/7.Man's impact on the natural vegetation in the Central Coastal Plain of Israel during the Chalcolithic and the Bronze Ages.Tel Aviv 13:71-84.

  3. Lev-Yadun, S. 1987.Dendrochronology and related studies - a review. Mitekufat Haeven, Journal of the Israel Prehistoric Society, n.s. 20:7-36.

  4. Liphschitz, N., S. Lev-Yadun & R. Gophna. 1987.The dominance of Quercus calliprinos Webb. (Kermes Oak) in the Central Coastal Plain of Israel in antiquity. Israel Exploration Journal 37:43-50.

  5. Liphschitz, N. & S. Lev-Yadun. 1989. The botanical remains from Masada. Identification of the plant species and the possible origin of the remnants.Bulletin of the American Schools of Oriental Research 274: 27-32.

  6. Lev-Yadun, S. 1992.The origin of the cedar beams from Al-Aqsa Mosque: Botanical, historical and archaeological evidence. Levant 24:201-208.

  7. Lev-Yadun, S. & R. Gophna. 1992.Exportation of plant products from Canaan to Egypt in the Early Bronze Age I: A rejoinder to William A. Ward.Bulletin of the American Schools of Oriental Research 287:89-90.

  8. Lev-Yadun, S. & M. Weinstein-Evron. 1993.Prehistoric wood remains of Cupressus sempervirens L. from the Natufian layers of el-Wad Cave, Mount Carmel, Israel.Tel Aviv 20:125-131.

  9. Goldberg, P., S. Lev-Yadun & O. Bar-Yosef. 1994.Petrographic thin sections of archaeological sediments: A new method for palaeobotanical studies.Geoarchaeology 9:243-257.

  10. Schiegl, S., S. Lev-Yadun, O. Bar-Yosef, A. El Goresy & S. Weiner. 1994.Siliceous aggregates from prehistoric wood ash: A major component of sediments in Kebara and Hayonim caves (Israel).Israel Journal of Earth Science 43:267-278.

  11. Lev-Yadun, S., Z. Herzog & T. Tsuk. 1995.Conifer beams of Juniperus phoenicea found in the well of Tel Beer-sheba.Tel Aviv 22:128-135.

  12. Lev-Yadun, S., Y. Mizrachi & M. Kochavi. 1996.Lichenometric studies of cultural formation processes at Rogem Hiri, Golan Heights.Israel Exploration Journal 46:196-207.

  13. Albert, R.M., O. Lavi, L. Estroff, S. Weiner, A. Tsatskin, A. Ronen & S. Lev-Yadun. 1999.Mode of occupation of Tabun Cave, Mt Carmel, Israel during the Mousterian period: A study of the sediments and phytoliths.Journal of Archaeological Sciences 26:1249-1260.

  14. Gopher, A., S. Abbo & S. Lev-Yadun. 2001.The “when”, the “where” and the “why” of the Neolithic Revolution in the Levant.Documenta Prehistorica (in press).

Papers in Books

  1. Lev-Yadun, S., N. Liphschitz & Y. Waisel. 1984. Ring analysis of Cedrus libani beams from the roof of al-Aqsa mosque.Eretz-Israel 17:92-96, 4*-5* (Hebrew and English* summary), The Israel Exploration Society, Jerusalem.

  2. Liphschitz, N., S. Lev-Yadun & Y. Waisel. 1987.A climatic history of the Sinai peninsula in light of dendrochronological studies. In: Sinai, Volume 1. Eds. Gvirtzman, G., A. Shmueli, Y. Gradus, I. Beit-Arieh & M. Har-El, pp. 525-531. Tel Aviv University, and Ministry of Defence Publishing House, Tel Aviv (Hebrew).

  3. Liphschitz, N., R. Gophna & S. Lev-Yadun. 1989.Man's impact on the vegetational landscape of Israel in the Early Bronze Age II-III. In: The urbanization of Palestine in the Early Bronze Age. Ed. Miroschedgji, P. de. BAR International Series, 527(ii), pp. 263-268.

  4. Lev-Yadun, S. 1997.Flora and climate in Southern Samaria: Past and Present. In: Highlands of many cultures I. The sites. Eds. Finkelstein, I. Lederman, Z. & S. Bunimovitz, pp. 85-102. Monograph Series of the Institute of Archaeology, Tel Aviv University, Tel Aviv.

  5. Lev-Yadun, S. 2000.Cellular patterns in dicotyledonous woods: their regulation.In: Cell & molecular biology of wood formation.Eds. Savidge, R., J. Barnett & R. Napier, pp. 315-324.BIOS Scientific Publishers Ltd., Oxford.

  6. Lev-Yadun, S. 2000.Wood structure and the ecology of annual growth ring formation in Pinus halepensis and Pinus brutia.In: Ecology, biogeography and management of Pinus halepensis and Pinus brutia forest ecosystems in the Mediterranean Basin.Eds. Ne`eman, G. & L. Trabaud, pp. 67-78.Backhuys Publishers, Leiden. (INCLUDING A COVER PICTURE).

  7. Weinstein-Evron, M. & S. Lev-Yadun. 2000.Paleoecology of Pinus halepensis in Israel in the light of archaeobotanical data.In: Ecology, biogeography and management of Pinus halepensis and Pinus brutia forest ecosystems in the Mediterranean Basin.Eds. Ne`eman, G. & L. Trabaud, pp. 119-130.Backhuys Publishers, Leiden. (INCLUDING A COVER PICTURE).

  8. Shmida, A., S. Lev-Yadun, S. Goubitz & G. Ne`eman. 2000.Sexual allocation and gender segregation in Pinus halepensis, P. brutia and P. pinea.In: Ecology, biogeography and management of Pinus halepensis and Pinus brutia forest ecosystems in the Mediterranean Basin.Eds. Ne`eman, G. & L. Trabaud, pp. 91-104.Backhuys Publishers, Leiden. (INCLUDING A COVER PICTURE).

  9. Lev-Yadun, S. 2001.The environmental influence of the Pottery Neolitic (ca. 7,500-6,500 years B.P.) Nahal Zehora II settlements in the southern Mt. Carmel.In: Pottery Neolithic Village in the southern Mt. Carmel: Nahal Zehora.Ed. Gopher, A.Monograph Series of the Institute of Archaeology, Tel Aviv University, Tel Aviv (in press).

  10. Lev-Yadun, S. 2002.Ecological aspects of the Late Bronze Age and Iron Age settlements of Tel Hadar.In: Tel Hadar I. The Early Iron Age pillared building complex.Eds. Beck, P., M. Kochavi & E. Yadin. Monograph Series of the Institute of Archaeology, Tel Aviv University, Tel Aviv (in press).

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