Isopoda oniscidea




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Malacostraca - Peracarida

ISOPODA - ONISCIDEA

LEILA APARECIDA SOUZA-KURY

Instituto de Biociências

Universidade de São Paulo

Suborder Oniscidea Latreille, 1803

Family Armadillidae Brandt & Ratzeburg, 1831

Genus Cubaris Brandt, 1833



Cubaris cinerea Brandt, 1833

Synonymy: van Name, 1936: 389.

Description and diagnosis: van Name, 1936: 389.

References: van Name, 1936: 389.

Geographic distribution: Brazil.
Cubaris murina Brandt, 1833

Synonymy: Green et al., 1990: 431.

Description and diagnosis: van Name, 1936: 387.

References: Lemos de Castro, 1967: 328; 1971: 12, fig. 13; 1972a: 357; Araújo et al., 1996: 132, figs. 49-53, 67.

Geographic distribution: Pantropical. Brazil (PA, BA, ES, MG, RJ, SP, SC).

Genus Diploexochus Brandt, 1833



Diploexochus echinatus Brandt, 1833

Synonymy: Lemos de Castro, 1967: 327.

Description and diagnosis: van Name, 1936: 398.

References: Lemos de Castro, 1967: 327.

Geographic distribution: Guyana, Trinidad, and Brazil (PA).

Genus Pseudodiploexochus Arcangeli, 1934



Pseudodiploexochus gibbus (Lemos de Castro, 1972)

Synonymy: Ferrara & Taiti, 1990: 489.

Description and diagnosis: Lemos de Castro, 1972b: 347.

References: Lemos de Castro, 1972b: 347, figs. 1-5.

Geographic distribution: Brazil (SP).

Comments: In moss.

Genus Venezillo Verhoeff, 1928



Venezillo congeneris (Budde-Lund, 1904)

Synonymy: van Name, 1936: 340.

Description and diagnosis: van Name, 1936: 340.

References: van Name, 1936: 340.

Geographic distribution: Brazil (MS).
Venezillo mineri (van Name, 1936)

Synonymy: van Name, 1936: 383.

Description and diagnosis: van Name, 1936: 383.

References: van Name, 1936: 383, fig. 233; Vandel, 1963: 94.

Geographic distribution: Guyana, Venezuela, and Brazil (PE, RJ).
Venezillo tuberosus (Budde-Lund, 1904)

Synonymy: van Name, 1936: 377.

Description and diagnosis: van Name, 1936: 377.

References: van Name, 1936: 377, fig. 228; Lemos de Castro, 1972a: 357.

Geographic distribution: Haiti, St. Thomas Island, and Brazil (BA).

Family Armadillidiidae Brandt, 1833

Genus Armadillidium Brandt, 1833

Armadillidium nasatum (Budde-Lund, 1885)

Synonymy: van Name, 1936: 279.

Description and diagnosis: Araújo et al., 1996: 136.

References: Araújo et al., 1996: 136, figs. 59-63, 68.

Geographic distribution: England, Italy, USA, and Brazil (RS)
Armadillidium vulgare (Latreille, 1804)

Synonymy: Vandel, 1962: 826.

Description and diagnosis: van Name, 1936: 276; Vandel, 1962: 826.

References: Camargo, 1954: 124; 1955: 2; Lemos de Castro, 1971: 8, fig. 14; Araújo et al., 1996: 133, figs. 54-58, 68.

Geographic distribution: Worldwide. Brazil (BA, MG, RJ, SP, SC, RS)

Family Balloniscidae Vandel, 1963

Genus Balloniscus Budde-Lund, 1908

Balloniscus glaber Araújo & Zardo, 1995

Synonymy: Araújo & Zardo, 1995: 785.

Description and diagnosis: Araújo & Zardo, 1995: 785.

References: Araújo & Zardo, 1995: 785, figs. 1-17.

Geographic distribution: Brazil (RS).

Balloniscus sellowi (Brandt, 1833)

Synonymy: Lemos de Castro, 1976: 392.

Description and diagnosis: Lemos de Castro, 1976: 392.

References: Lemos de Castro, 1972a: 357; 1976: 392, figs. 1-13; Araújo et al., 1996: 120, figs. 22-27, 65.

Geographic distribution: Brazil (BA, ES, MG, RJ, SP, PR, SC, RS), Paraguay, Uruguay, and Argentina.

Family Bathytropidae Vandel, 1952

Genus Neotroponiscus Arcangeli, 1936

Neotroponiscus argentinus (Giambiagi de Calabrese, 1939)

Synonymy: Lemos de Castro, 1970b: 93.

Description and diagnosis: Andersson, 1960: 560; Lemos de Castro, 1970b: 93.

References: Vandel, 1963: 82; Lemos de Castro, 1970b: 93, fig 3.

Geographic distribution: Argentina and Brazil (PE, ES, RJ, SP, SC).

Comments: Under logs and stones; beneath sheaths of banana trees and bromeliads.
Neotroponiscus carolii Arcangeli, 1936

Synonymy: Lemos de Castro, 1970b: 90.

Description and diagnosis: Lemos de Castro, 1970b: 90.

References: Lemos de Castro, 1970b: 90, figs. 1-2.

Geographic distribution: Brazil (BA, ES, SP).

Comments: Myrmecophilous in nests of Camponotus rufipes; under rotten logs.

Neotroponiscus daguerrii (Giambiagi de Calabrese, 1939)

Synonymy: Araújo et al., 1996: 122.

Description and diagnosis: Araújo et al., 1996: 122.

References: Araújo et al., 1996: 122, figs. 28-38, 65.

Geographic distribution: Brazil (RS) and Argentina.

Neotroponiscus lenkoi Lemos de Castro, 1970

Synonymy: Lemos de Castro, 1970c: 8.

Description and diagnosis: Lemos de Castro, 1970c: 8.

References: Lemos de Castro, 1970c: 8, figs. 5-6.

Geographic distribution: Brazil (SP).
Neotroponiscus littoralis Lemos de Castro, 1970

Synonymy: Lemos de Castro, 1970c: 1.

Description and diagnosis: Lemos de Castro, 1970c: 1.

References: Lemos de Castro, 1970c: 1, figs. 1-2.

Geographic distribution: Brazil (RJ).

Neotroponiscus lobatus Lemos de Castro, 1970

Synonymy: Lemos de Castro, 1970c: 5.

Description and diagnosis: Lemos de Castro, 1970c: 5.

References: Lemos de Castro, 1970c: 5, figs. 3-4.

Geographic distribution: Brazil (ES).

Neotroponiscus perlatus Lemos de Castro, 1970

Synonymy: Lemos de Castro, 1970c: 11.

Description and diagnosis: Lemos de Castro, 1970c: 11.

References: Lemos de Castro, 1970c: 11, figs. 7-8.

Geographic distribution: Brazil (ES).
Neotroponiscus plaumanni (Andersson, 1960)

Synonymy: Lemos de Castro, 1970b: 93.

Description and diagnosis: Andersson, 1960: 563; Lemos de Castro, 1970b: 93.

References: Lemos de Castro, 1970b: 93, fig. 4.

Geographic distribution: Brazil (SC) and Uruguay.

Family Dubioniscidae Schultz, 1995

Genus Calycuoniscus Collinge, 1915

Calycuoniscus bodkini Collinge, 1915

Synonymy: Lemos de Castro, 1968: 410.

Description and diagnosis: Lemos de Castro, 1968: 410.

References: Lemos de Castro, 1968: 410, figs. 8-16.

Geographic distribution: Trinidad, Guyana, and Brazil (AP, PA).

Comments: Under bark. In nests of guacharos in cave. Under pieces of wood on river banks.
Calycuoniscus goeldii (Lemos de Castro, 1967)

Synonymy: Lemos de Castro, 1968: 408.

Description and diagnosis: Lemos de Castro, 1967: 318; 1968: 408.

References: Lemos de Castro, 1967: 318; 1968: 410, figs. 1-7.

Geographic distribution: Brazil (PA).

Comments: Under debris on moist soil. In layers of rotten fibers and debris in palm trunk 50 cm above the ground.

Genus Dubioniscus Vandel, 1963



Dubioniscus delamarei Vandel, 1963

Synonymy: Schultz, 1995: 401.

Description and diagnosis: Vandel, 1963: 78; Schultz, 1995: 401.

References: Lemos de Castro, 1970a: 2; Schultz, 1995: 401, figs. 5-7.

Geographic distribution: Brazil (ES, MG, RJ, SP), Paraguay, and Argentina.

Comments: Myrmecophilous.
Dubioniscus marmoratus Lemos de Castro, 1970

Synonymy: Lemos de Castro, 1970a: 3.

Description and diagnosis: Lemos de Castro, 1970a: 3.

References: Lemos de Castro, 1970a: 3, figs. 1-3.

Geographic distribution: Brazil (RJ, SP).

Genus Novamundoniscus Schultz, 1995



Novamundoniscus dissimilis (Lemos de Castro, 1959)

Synonymy: Lemos de Castro, 1959: 207; Schultz, 1995: 407.

Description and diagnosis: Lemos de Castro, 1959: 207.

References: Lemos de Castro, 1959: 207, figs. 28-35.

Geographic distribution: Brazil (RJ).
Novamundoniscus macrophthalmus (Lemos de Castro, 1959)

Synonymy: Lemos de Castro, 1959: 205; Schultz, 1995: 407.

Description and diagnosis: Lemos de Castro, 1959: 205.

References: Lemos de Castro, 1959: 205, figs. 10-18.

Geographic distribution: Brazil (RJ).
Novamundoniscus persimilis (Vandel, 1952)

Synonymy: Lemos de Castro, 1967: 322; Schultz, 1995: 407.

Description and diagnosis: Vandel, 1952a: 144; Lemos de Castro, 1967: 322.

References: Lemos de Castro, 1967: 322.

Geographic distribution: Venezuela and Brazil (PA).
Novamundoniscus singularis (Lemos de Castro, 1967)

Synonymy: Lemos de Castro, 1970d: 119; Schultz, 1995: 407.

Description and diagnosis: Lemos de Castro, 1967: 321; 1970d: 119.

References: Lemos de Castro, 1967: 321; 1970d: 119, figs. 1-11.

Geographic distribution: Brazil (AM).
Novamundoniscus vandeli (Lemos de Castro, 1959)

Synonymy: Lemos de Castro, 1959: 205; Schultz, 1995: 407.

Description and diagnosis: Lemos de Castro, 1959: 205.

References: Lemos de Castro, 1959: 205, figs. 1-9.

Geographic distribution: Brazil (MG, RJ) and Paraguay.

Genus Phalloniscus Budde-Lund, 1908



Phalloniscus loyolai Zardo, 1989

Synonymy: Zardo, 1989: 611.

Description and diagnosis: Zardo, 1989: 611.

References: Zardo, 1989: 611, figs 1-25.

Geographic distribution: Brazil (PR).
Phalloniscus meridionalis Araújo & Buckup, 1994

Synonymy: Araújo & Buckup, 1994a: 271.

Description and diagnosis: Araújo & Buckup, 1994a: 271.

References: Araújo & Buckup, 1994a: 271, figs. 14-28.

Geographic distribution: Brazil (SC, RS).
Phalloniscus setosus Lemos de Castro, 1959

Synonymy: Lemos de Castro, 1959: 207.

Description and diagnosis: Lemos de Castro, 1959: 207.

References: Lemos de Castro, 1959: 207, figs 19-27.

Geographic distribution: Brazil (MG).

Family Eubelidae Budde-Lund, 1899

Genus Ethelum Budde-Lund, 1899

Ethelum americanum (Dollfus, 1896)

Synonymy: Lemos de Castro, 1967: 327.

Description and diagnosis: van Name, 1927: 484.

References: Lemos de Castro, 1967: 327.

Geographic distribution: West Indies, Guyana, French Guyana, and Brazil (PA).

Family Ligiidae Brandt & Ratzeburg, 1831

Genus Ligia Fabricius, 1798

Ligia baudiniana Milne-Edwards, 1840

Synonymy: Schultz, 1984: 4.

Description and diagnosis: van Name, 1936: 58.

References: van Name, 1936: 58, fig. 14; Vandel, 1952a: 80.

Geographic distribution:, United States, Galapagos, Mexico, Venezuela, West Indies, and Brazil (PB, PE, RJ).

Comments: Halophilous; under stones and on moist rocks near water line; estuaries.

Ligia exotica Roux, 1828

Synonymy: Ferrara & Taiti, 1979: 93.

Description and diagnosis: Chilton, 1916: 463; Brian & Dartevelle, 1949: 95; Roman, 1977: 119.

References: van Name, 1936; Andersson, 1960: 540.

Geographic distribution: Pantropical. Brazil (RJ).

Comments: Halophilous.

Ligia olfersii Brandt, 1833

Synonymy: van Name, 1936: 53.

Description and diagnosis: van Name, 1936: 53.

References: Moreira, 1932: 433; van Name, 1936: 53, fig. 11.

Geographic distribution: United States, West Indies, Brazil (RJ), and western and central Africa.

Comments: Halophilous.

Genus Stymphalus Budde-Lund, 1885



Stymphalus dilatatus (Perty, 1833)V

Synonymy: van Name, 1936: 66.

Description and diagnosis: van Name, 1936: 66.

References: van Name, 1936: 66, fig. 21.

Geographic distribution: Brazil ("near BA").

Family Olibrinidae Budde-Lund, 1913

Genus Olibrinus Budde-Lund, 1913

Olibrinus sp.

Synonymy: Lemos de Castro, 1972a: 357.

Description and diagnosis: Lemos de Castro, 1972a: 357.

References: Lemos de Castro, 1972a: 357.

Geographic distribution: Brazil (BA).

Family Oniscidae Latreille, 1806

Genus Oniscus Linnaeus, 1758

Oniscus asellus Linnaeus, 1758

Synonymy: Vandel, 1962: 538.

Description and diagnosis: van Name, 1936: 182; Vandel, 1962: 538.

References: Zardo & Loyola e Silva, 1988: 791, figs. 1-4.

Geographic distribution: Europe, Canada, USA, Cuba, St Helen Island, Azores, Madeira, and Brazil (PR) .

Comments: Western European origin, introduced in the Americas.

Family Philosciidae Vandel, 1952; Taiti & Ferrara, 1980: 54; 1982: 1

Genus Atlantoscia Ferrara & Taiti, 1981

Atlantoscia floridana (van Name, 1940)

Synonymy: Taiti & Ferrara, 1991a: 902.

Description and diagnosis: Taiti & Ferrara, 1991a: 902.

References: Lemos de Castro, 1985b: 418, figs. 1-15; Taiti & Ferrara, 1991a: 902, figs. 1-15; Araújo et al., 1996: 115, figs. 11-14, 64.

Geographic distribution: United States, Brazil (AP, PA, RN, PB, PE, BA, ES, MG, RJ, SP, PR, SC, RS, TI), Argentina, Ascension and St. Helena Islands.

Genus Benthana Budde-Lund, 1908



Benthana albomarginata Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 95.

Description and diagnosis: Lemos de Castro, 1958a: 95.

References: Lemos de Castro, 1958a: 95, figs. 16-27.

Geographic distribution: Brazil (ES).

Comments: In epiphytic bromeliads.
Benthana bocainensis Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 113.

Description and diagnosis: Lemos de Castro, 1958a: 113.

References: Lemos de Castro, 1958a: 113, figs. 87-92.

Geographic distribution: Brazil (SP).

Benthana convexa Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 93.

Description and diagnosis: Lemos de Castro, 1958a: 93.

References: Lemos de Castro, 1958a: 93, figs. 10-15.

Geographic distribution: Brazil (SP).

Benthana dimorpha Lemos de Castro, 1985

Synonymy: Lemos de Castro, 1985a: 246.

Description and diagnosis: Lemos de Castro, 1985a: 246.

References: Lemos de Castro, 1985a: 246, figs. 14-25.

Geographic distribution: Brazil (ES).

Benthana iporangensis Lima & Serejo, 1993

Synonymy: Lima & Serejo, 1993: 490.

Description and diagnosis: Lima & Serejo, 1993: 490.

References: Lima & Serejo, 1993: 490, figs. 1-4.

Geographic distribution: Brazil (SP).

Comments: Cave-dwelling, troglomorphic.

Benthana longicornis Verhoeff, 1941

Synonymy: Lemos de Castro, 1958a: 115.

Description and diagnosis: Lemos de Castro, 1958a: 115.

References: Lemos de Castro, 1958a: 115, figs. 93-98; Andersson, 1960: 557, fig. 10; Araújo et al., 1996: 115, figs. 5-10, 64.

Geographic distribution: Brazil (SC).

Comments: Under logs on river banks; under stones and pieces of wood.
Benthana longipenis Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 109.

Description and diagnosis: Lemos de Castro, 1958a: 109.

References: Lemos de Castro, 1958a: 109, figs. 69-77.

Geographic distribution: Brazil (SP).

Comments: Under bark, rotten logs and in bromeliads in coppices.
Benthana moreirai Lemos de Castro, 1985

Synonymy: Lemos de Castro, 1985a: 241.

Description and diagnosis: Lemos de Castro, 1985a: 241.

References: Lemos de Castro, 1985a: 241, figs. 1-13.

Geographic distribution: Brazil (SP).

Comments: Under sticks and leaves.
Benthana olfersii (Brandt, 1833)

Synonymy: Lemos de Castro, 1958a: 98.

Description and diagnosis: Lemos de Castro, 1958a: 98.

References: Lemos de Castro, 1958a: 98, figs. 28-39.

Geographic distribution: Germany (in orchids brought from southern Brazil) and Brazil (RJ, SP).

Comments: Under stones and leaves in woods.

Benthana picta (Brandt, 1833)

Synonymy: Lemos de Castro, 1958a: 111.

Description and diagnosis: Lemos de Castro, 1958a: 111.

References: Lemos de Castro, 1958a: 111, figs 78-86; Araújo et al., 1996: 113, figs. 1-4, 64.

Geographic distribution: Germany (in plants brought from Brazil), Brazil (RJ, SP, SC, RS), and Argentina.
Benthana santosi Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 106.

Description and diagnosis: Lemos de Castro, 1958a: 106.

References: Lemos de Castro, 1958a: 106, figs. 60-68.

Geographic distribution: Brazil (MG, RJ, SP).
Benthana schubarti Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 91.

Description and diagnosis: Lemos de Castro, 1958a: 91.

References: Lemos de Castro, 1958a: 91, figs. 1-9.

Geographic distribution: Brazil (RJ, SP).
Benthana sulcata Gruner, 1955

Synonymy: Lemos de Castro, 1958a: 102.

Description and diagnosis: Lemos de Castro, 1958a: 102.

References: Lemos de Castro, 1958a: 102, figs. 40-49.

Geographic distribution: Brazil (RJ).

Comments: In bromeliads.
Benthana taeniata Araújo & Buckup, 1994

Synonymy: Araújo & Buckup, 1994b: 269.

Description and diagnosis: Araújo & Buckup, 1994b: 269.

References: Araújo & Buckup, 1994b: 269, figs. 1-13, 28.

Geographic distribution: Brazil (SC, RS).
Benthana werneri Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958a: 104.

Description and diagnosis: Lemos de Castro, 1958a: 104.

References: Lemos de Castro, 1958a: 104, figs. 50-59.

Geographic distribution: Brazil (SP).

Genus Benthanoscia Lemos de Castro, 1958



Benthanoscia longicaudata Lemos de Castro, 1958

Synonymy: Lemos de Castro, 1958b: 2.

Description and diagnosis: Lemos de Castro, 1958b: 2.

References: Lemos de Castro, 1958b: 2, figs. 1-14.

Geographic distribution: Brazil (RJ).

Genus Burmoniscus Collinge, 1914



Burmoniscus meeusei (Holthuis, 1947)

Synonymy: Taiti & Ferrara, 1991b: 212.

Description and diagnosis: Araújo et al., 1996: 118.

References: Araújo et al., 1996: 118, figs. 15-21, 64.

Geographic distribution: England (hothouses), Hawaii, and Brazil (SC).

Genus Chaetophiloscia Verhoeff, 1908



Chaetophiloscia frontalis Lemos de Castro, 1967

Synonymy: Lemos de Castro, 1967: 320.

Description and diagnosis: Lemos de Castro, 1967: 320.

References: Lemos de Castro, 1967: 320.

Geographic distribution: Brazil (PA).
Chaetophiloscia gatunensis (van Name, 1926)

Synonymy: Lemos de Castro, 1967: 321 .

Description and diagnosis: van Name, 1936: 158.

References: Lemos de Castro, 1967: 321.

Geographic distribution: Panama and Brazil (AM, PA).
Chaetophiloscia walkeri (Pearse, 1915)

Synonymy: Lemos de Castro, 1967: 319.

Description and diagnosis: van Name, 1936: 153.

References: Lemos de Castro, 1967: 319.

Geographic distribution: Colombia and Brazil (PA).

Genus Ischioscia Verhoeff, 1928



Ischioscia amazonica Lemos de Castro, 1955

Synonymy: Schmalfuß, 1980: 131.

Description and diagnosis: Lemos de Castro, 1955: 51; Vandel, 1968: 78.

References: Lemos de Castro, 1955: 51, figs. 1-8; 1967: 318.

Geographic distribution: Ecuador and Brazil (AM).

Comments: Under bark of dead trees in transitional forest.
Ischioscia irmleri Schmalfuß, 1980

Synonymy: Schmalfuß, 1980: 136.

Description and diagnosis: Schmalfuß, 1980: 136.

References: Schmalfuß, 1980: 136, figs. 27-31.

Geographic distribution: Brazil (AM).

Comments: Collected with traps in “igapó”.
Genus Littorophiloscia Hatch, 1947

Littorophiloscia tropicalis Taiti & Ferrara, 1986

Synonymy: Taiti & Ferrara, 1986: 1361.

Description and diagnosis: Schultz, 1983: 441; Taiti & Ferrara, 1986: 1361.

References: Lemos de Castro, 1968: 94, figs. 31-33.

Geographic distribution: India, Cameroon, Somalia, Sudan, United States, Mexico, Belize, Venezuela, and Brazil (RJ).

Genus Parischioscia Lemos de Castro, 1967



Parischioscia omissa (van Name, 1936)

Synonymy: Lemos de Castro, 1967: 319.

Description and diagnosis: van Name, 1936: 140.

References: Lemos de Castro, 1967: 319.

Geographic distribution: Guyana, French Guyana, and Brazil (AP).

Genus Prosekia Vandel, 1968



Prosekia albamaculata Lima, 1996

Synonymy: Lima, 1996a: 102.

Description and diagnosis: Lima, 1996a: 102

References: Lima, 1996a: 102, figs. 1-17.

Geographic distribution: Brazil (AM).
Prosekia insularis Lemos de Castro & Souza, 1986

Synonymy: Lemos de Castro & Souza, 1986: 435.

Description and diagnosis: Lemos de Castro & Souza, 1986: 435.

References: Lemos de Castro & Souza, 1986: 435, figs. 27-41.

Geographic distribution: Brazil (PA).
Prosekia lejeunei Lemos de Castro & Souza, 1986

Synonymy: Lemos de Castro & Souza, 1986: 432.

Description and diagnosis: Lemos de Castro & Souza, 1986: 432.

References: Lemos de Castro & Souza, 1986: 432, figs. 13-26.

Geographic distribution: Brazil (PA).
Prosekia silvatica Lemos de Castro & Souza, 1986

Synonymy: Lemos de Castro & Souza, 1986: 432.

Description and diagnosis: Lemos de Castro & Souza, 1986: 432.

References: Lemos de Castro & Souza, 1986: 432, figs. 1-12.

Geographic distribution: Brazil (AM, PA).

Comments: On bark of fallen log; under vegetable debris in forest.
Prosekia tarumae Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984b: 442.

Description and diagnosis: Lemos de Castro, 1984b: 442.

References: Lemos de Castro, 1984b: 442, figs. 1-12.

Geographic distribution: Brazil (AM).

Family Platyarthridae Verhoeff, 1949

Genus Niambia Budde-Lund, 1909

Niambia squamata (Budde-Lund, 1885)

Synonymy: Schmalfuß & Ferrara, 1978: 77.

Description and diagnosis: Brian, 1953: 11.

References: Lemos de Castro, 1967: 315; 1971: 3; 1972a: 357.

Geographic distribution: Southern Africa and Brazil (PA, BA, FN).

Genus Trichorhina Budde-Lund, 1908



Trichorhina acuta Araújo & Buckup, 1994

Synonymy: Araújo & Buckup, 1994b: 130.

Description and diagnosis: Araújo & Buckup, 1994b: 130.

References: Araújo & Buckup, 1994b: 130, figs. 1-12.

Geographic distribution: Brazil (SC, RS).
Trichorhina amazonica Souza-Kury, 1997

Synonymy: Souza-Kury, 1997a: 183.

Description and diagnosis: Souza-Kury, 1997a: 183.

References: Souza-Kury, 1997a: 183, figs. 2, 8-20.

Geographic distribution: Brazil (PA).
Trichorhina brasilensis Andersson, 1960

Synonymy: Schultz, 1995: 413.

Description and diagnosis: Andersson, 1960: 552; Schultz, 1995: 413.

References: Andersson, 1960: 552, fig. 8.

Geographic distribution: Brazil (SC) and Paraguay.

Comments: Under stones.
Trichorhina guanophila Souza-Kury, 1993

Synonymy: Souza-Kury, 1993a: 198.

Description and diagnosis: Souza-Kury, 1993a: 198.

References: Souza-Kury, 1993a: 198, figs. 11-27.

Geographic distribution: Brazil (BA).

Comments: In bat guano in cave.
Trichorhina heterophthalma Lemos de Castro, 1964

Synonymy: Souza-Kury, 1993a: 198.

Description and diagnosis: Lemos de Castro, 1964: 2; Souza-Kury, 1993a: 198.

References: Souza-Kury, 1993a: 198, figs. 1-8.

Geographic distribution: Reunion, Galapagos, United States, Mexico, West Indies, Cuba, Brazil (RJ), and Paraguay.

Trichorhina macrops Souza-Kury, 1993

Synonymy: Souza-Kury, 1993a: 205.

Description and diagnosis: Souza-Kury, 1993a: 205.

References: Souza-Kury, 1993a: 205, figs. 28-39.

Geographic distribution: Brazil (PE).

Trichorhina paraensis Souza-Kury, 1997

Synonymy: Souza-Kury, 1997a: 186.

Description and diagnosis: Souza-Kury, 1997a: 186.

References: Souza-Kury, 1997a: 186, figs. 3, 21-36.

Geographic distribution: Brazil (PA).
Trichorhina pittieri (Pearse, 1921)

Synonymy: van Name, 1936: 200.

Description and diagnosis: van Name, 1936: 200.

References: Lemos de Castro, 1967: 316.

Geographic distribution: Venezuela, Guyana, and Brazil (PA).
Trichorhina tomentosa (Budde-Lund, 1893)

Synonymy: Souza-Kury, 1991: 23.

Description and diagnosis: Souza-Kury, 1991: 23.

References: Vandel, 1963: 72. Lemos de Castro, 1967: 315; 1971: 10, fig. 6; Souza-Kury, 1991: 23, figs. 14-19.

Geographic distribution: Worldwide. Brazil (PA, ES, MG, RJ, SP).

Family Porcellionidae Brandt & Ratzeburg, 1831; Vandel, 1962: 575

Genus Agabiformius Verhoeff, 1908

Agabiformius lentus (Budde-Lund, 1885)

Synonymy: Vandel, 1962: 638.

Description and diagnosis: Vandel, 1962: 638.

References: Lemos de Castro, 1971: 7, fig. 8.

Geographic distribution: Worldwide. Brazil (RJ).

Comments: Mediterranean origin, introduced in many continents.

Genus Porcellio Latreille, 1804



Porcellio dilatatus Brandt, 1833

Synonymy: Vandel, 1962: 672.

Description and diagnosis: Vandel, 1962: 672.

References: Lemos de Castro, 1971: 5, fig. 11; Araújo et al., 1996: 126, figs. 41, 42, 66.

Geographic distribution: Europe (Portugal to Russia, and Turkey) and Brazil (MG, RJ, SP, PR, SC, RS).

Comments: Iberic origin, introduced in the Americas, Europe up to Russia, and Turkey.
Porcellio laevis Latreille, 1804

Synonymy: Vandel, 1962: 684.

Description and diagnosis: Vandel, 1962: 684.

References: Moreira, 1927: 194; Lemos de Castro, 1971: 4, fig. 10.

Geographic distribution: Worldwide. Brazil (RJ, SP).
Porcellio scaber Latreille, 1804

Synonymy: Ferrari & Taiti, 1979: 130.

Description and diagnosis: Araújo et al., 1996: 125.

References: Araújo et al., 1996: 125.

Geographic distribution: Europe, Africa, St Helen Island, Ascencion Island, Tristan da Cunha Island, and South America. Brazil (SC, RS).

Comments: Western European origin, introduced in Africa and Americas.

Genus Porcellionides Miers, 1877



Porcellionides advena (Stuxberg, 1872)

Synonymy: van Name, 1936: 247.

Description and diagnosis: van Name, 1936: 247.

References: van Name, 1936: 247, figs. 139-140.

Geographic distribution: Brazil (MG) .
Porcellionides pruinosus (Brandt, 1833)

Synonymy: Vandel, 1962: 618.

Description and diagnosis: Vandel, 1962: 618; Lemos de Castro, 1971: 5.

References: Lemos de Castro, 1967: 323; 1972a: 357; Araújo et al., 1996: 129, figs. 45-47.

Geographic distribution: Worldwide. Brazil (PA, BA, SC, RS).

Comments: Under stones, near houses.
Porcellionides schwencki (Moreira, 1927)

Synonymy: van Name, 1936: 238.

Description and diagnosis: Moreira, 1927: 194; Giambiagi de Calabrese, 1939: 638.

References: Moreira, 1927: 194, figs. 1-3.

Geographic distribution: Brazil (SP) and Argentina.

Comments: In garden earth, under stones, pots and other objects. Considered as possible synonym of P. pruinosus (Lemos de Castro, 1971: 6). Giambiagi de Calabrese (1939: 639) considered that there are enough diagnostic characters to separate both species.
Porcellionides sexfasciatus (Koch, 1847)

Synonymy: Zardo & Loyola e Silva, 1988: 791.

Description and diagnosis: Araújo et al., 1996: 130.

References: Araújo et al., 1996: 130, figs. 47-48, 67.

Geographic distribution: Worldwide. Brazil (PR, SC, RS).
Family Pudeoniscidae Lemos de Castro, 1973

Genus Braziloniscus Lemos de Castro, 1973



Braziloniscus maculatus Lemos de Castro, 1973

Synonymy: Lemos de Castro, 1973: 6.

Description and diagnosis: Lemos de Castro, 1973: 6.

References: Lemos de Castro, 1973: 6, figs. 3-4.

Geographic distribution: Brazil (RJ, SP).
Braziloniscus verrucosus Lemos de Castro, 1973

Synonymy: Lemos de Castro, 1973: 9.

Description and diagnosis: Lemos de Castro, 1973: 9.

References: Lemos de Castro, 1973: 9, fig. 5.

Geographic distribution: Brazil (RJ).

Genus Pudeoniscus Vandel, 1963



Pudeoniscus birabeni Vandel, 1963

Synonymy: Lemos de Castro, 1973: 3 .

Description and diagnosis: Vandel, 1963: 91.

References: Vandel, 1963: 91, figs. 16-19; Lemos de Castro, 1973: 3.

Geographic distribution: Brazil (RJ, SP).
Pudeoniscus obscurus Lemos de Castro, 1973

Synonymy: Lemos de Castro, 1973: 4.

Description and diagnosis: Lemos de Castro, 1973: 4.

References: Lemos de Castro, 1973: 4, figs. 1-2.

Geographic distribution: Brazil (SP).

Family Rhyscotidae Budde-Lund, 1904

Genus Rhyscotus Budde-Lund, 1885

Rhyscotus albidemaculatus Budde-Lund, 1908

Synonymy: Souza-Kury, 1997b: 106.V

Description and diagnosis: Souza-Kury, 1997b: 106.

References: Souza-Kury, 1997b: 106, figs. 1-10.

Geographic distribution: Brazil (BA, RJ).

Family Scleropactidae Verhoeff, 1938

Genus Amazoniscus Lemos de Castro, 1967

Amazoniscus arlei Lemos de Castro, 1967

Synonymy: Lemos de Castro, 1969: 2.

Description and diagnosis: Lemos de Castro, 1967: 326; 1969: 2.

References: Lemos de Castro, 1967: 326; 1969: 2, figs 1-11.

Geographic distribution: Brazil (AP, PA).

Genus Circoniscus Pearse, 1917



Circoniscus amazonicus Lima, 1996

Synonymy: Lima, 1996b: 92.

Description and diagnosis: Lima, 1996b: 92.

References: Lima, 1996b: 92, figs. 1-26.

Geographic distribution: Brazil (AM).
Circoniscus bezzii Arcangeli, 1931

Synonymy: Souza & Lemos de Castro, 1991: 50; Schultz, 1995: 417.

Description and diagnosis: Souza & Lemos de Castro, 1991: 50.

References: Souza & Lemos de Castro, 1991: 50, figs. 23-44.

Geographic distribution: Brazil (MT, ES, MG, SP) and Paraguay.
Circoniscus gaigei Pearse, 1917

Synonymy: Souza & Lemos de Castro, 1991: 47.

Description and diagnosis: Souza & Lemos de Castro, 1991: 47.

References: Lemos de Castro, 1967: 324; Souza & Lemos de Castro, 1991: 47, figs. 1-22.

Geographic distribution: Peru, Guyana, French Guyana, and Brazil (AM, AP, PA).
Circoniscus gracilidens Souza & Lemos de Castro, 1991

Synonymy: Souza & Lemos de Castro, 1991: 59.

Description and diagnosis: Souza & Lemos de Castro, 1991: 59.

References: Souza & Lemos de Castro, 1991: 59, figs. 91-107.

Geographic distribution: Brazil (RJ).
Circoniscus incisus Souza & Lemos de Castro, 1991

Synonymy: Souza & Lemos de Castro, 1991: 56.

Description and diagnosis: Souza & Lemos de Castro, 1991: 56.

References: Souza & Lemos de Castro, 1991: 56, figs. 69-90.

Geographic distribution: Brazil (RJ).
Circoniscus intermedius Souza & Lemos de Castro, 1991

Synonymy: Souza & Lemos de Castro, 1991: 53.

Description and diagnosis: Souza & Lemos de Castro, 1991: 53.

References: Souza & Lemos de Castro, 1991: 53, figs. 45-68.

Geographic distribution: Brazil (PA, MT).

Genus Microsphaeroniscus Lemos de Castro, 1984



Microsphaeroniscus bicolor Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984a: 3.

Description and diagnosis: Lemos de Castro, 1984a: 3.

References: Lemos de Castro, 1984a: 3, figs. 18-21.

Geographic distribution: Brazil (SP).

Microsphaeroniscus costatus Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984a: 3.

Description and diagnosis: Lemos de Castro, 1984a: 3.

References: Lemos de Castro, 1984a: 3, figs. 22-26.

Geographic distribution: Brazil (RJ).
Microsphaeroniscus pallidus Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984a: 2.

Description and diagnosis: Lemos de Castro, 1984a: 2.

References: Lemos de Castro, 1984a: 2, figs. 1-14.

Geographic distribution: Brazil (RJ, SP)
Microsphaeroniscus squamatus Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984a: 4.

Description and diagnosis: Lemos de Castro, 1984a: 4.

References: Lemos de Castro, 1984a: 4, figs. 27-38.

Geographic distribution: Brazil (RJ).
Microsphaeroniscus violaceus Lemos de Castro, 1984

Synonymy: Lemos de Castro, 1984a: 3.

Description and diagnosis: Lemos de Castro, 1984a: 3.

References: Lemos de Castro, 1984a: 3, figs. 15-17.

Geographic distribution: Brazil (SP).

Genus Parsphaeroniscus Vandel, 1963



Parsphaeroniscus apeuensis Lemos de Castro, 1967

Synonymy: Lemos de Castro, 1967: 325.

Description and diagnosis: Lemos de Castro, 1970e: 41.

References: Lemos de Castro, 1970e: 41, figs. 1-6.

Geographic distribution: Brazil (PA).

Family Styloniscidae Vandel, 1952

Genus Clavigeroniscus Arcangeli, 1930

Clavigeroniscus riquieri (Arcangeli, 1930)

Synonymy: Lemos de Castro, 1967: 314.

Description and diagnosis: Vandel, 1952b: 70.

References: Lemos de Castro, 1967: 314.

Geographic distribution: Africa, Central and South America. Brazil (AP, PA).

Genus Cordioniscus Graeve, 1914



Cordioniscus stebbingi (Patience, 1907)

Synonymy: Lemos de Castro, 1953: 533.

Description and diagnosis: Vandel, 1952b: 63; Lemos de Castro, 1953: 533.

References: Lemos de Castro, 1953: 533, figs. 14-22.; 1971: 2, fig. 1.

Geographic distribution: England, United States, and Brazil (RJ, SP).

Genus Pectenoniscus Andersson, 1960



Pectenoniscus angulatus Andersson, 1960

Synonymy: Andersson, 1960: 550.

Description and diagnosis: Andersson, 1960: 550.

References: Andersson, 1960: 550, figs. 6-7.

Geographic distribution: Brazil (SC).

Family Trachelipodidae Strouhal, 1953



Comments: The name of this family has been recently corrected (Souza-Kury, 1993b).

Genus Nagurus Holthuis, 1949



Nagurus cristatus (Dollfus, 1889)

Synonymy: Green et al., 1990: 429.

Description and diagnosis: Lemos de Castro, 1971: 7.

References: Lemos de Castro, 1967: 323; 1971: 7, fig. 7.

Geographic distribution: Pantropical. Brazil (PA, RJ, SP).

Genus Trachelipus Budde-Lund, 1908



Trachelipus rathkei (Brandt, 1833)

Synonymy: Vandel, 1962: 584.

Description and diagnosis: Vandel, 1962: 584.

References: Lemos de Castro, 1971: 6, fig. 12.

Geographic distribution: Europe, Canada, United States, Mexico, Brazil (RJ), and Argentina.

Family Trichoniscidae Sars, 1899

Genus Haplophthalmus Schöbl, 1899

Haplophthalmus danicus Budde-Lund, 1879

Synonymy: Vandel, 1977: 387.

Description and diagnosis: Lemos de Castro, 1971: 3.

References: Lemos de Castro, 1971: 3, fig. 2.

Geographic distribution: Japan, Asia Minor, Europe, Madeira, Canaries, Azores, northern Africa, North America, and Brazil (SP).

Genus Miktoniscus Kesselyak, 1930



Miktoniscus medcofi (van Name, 1940)

Synonymy: Lemos de Castro, 1953: 529.

Description and diagnosis: Lemos de Castro, 1953: 529.

References: Lemos de Castro, 1953: 529, figs. 1-13; 1971: 10, fig. 3.

Geographic distribution: United States and Brazil (PA, RJ, SP).

Comments: In banana growth. Only greenhouses in USA.

Family Tylidae Milne Edwards, 1840

Genus Tylos Audouin, 1826

Tylos niveus Budde-Lund, 1885

Synonymy: Mulaik, 1960: 93.

Description and diagnosis: Mulaik, 1960: 93.

References: Lemos de Castro, 1952: 2, figs. 1-15.

Geographic distribution: United States, Mexico, Bermudas, Cuba, and Brazil (RJ).

Comments: According to OPINION 369 of ICZN (1955), authorship of genus Tylos should be Audouin, not Latreille, and date 1826, not 1825.

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Zardo, C.M.L. & Loyola e Silva, J. 1988. Primeira ocorrência de Oniscus asellus Linné, 1758 e Porcellionides sexfasciatus (Koch, 1847) (Isopoda, Oniscoidea) no Brasil. Ciência e Cultura 40(8): 791-795.


 SOUZA-KURY, L.A., 1998. Malacostraca - Peracarida. Isopoda. Oniscidea. In: YOUNG, P.S. (ed.). Catalogue of Crustacea of Brazil. Rio de Janeiro: Museu Nacional. p. 653-674. (Série Livros n. 6).



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