Including Canada, the continental United States, and the northern Mexican states of Baja California, Baja California Sur, Chihuahua, Coahuila de Zaragoza, Durango, Nuevo Leon, Sinaloa, Sonora, Tamaulipas, and Zacatecas




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Checklist and Nomenclatural Authority File of the Scarabaeoidea of the Nearctic Realm

including Canada, the continental United States, and the northern Mexican states of Baja California, Baja California Sur, Chihuahua, Coahuila de Zaragoza, Durango, Nuevo Leon, Sinaloa, Sonora, Tamaulipas, and Zacatecas.

Version 4 - released 22 April 2009




Andrew B. T. Smith



Canadian Museum of Nature

P.O. Box 3443, Station D

Ottawa, ON, K1P 6P4

CANADA

asmith@mus-nature.ca

SCARABAEOIDEA Latreille, 1802

PLEOCOMIDAE LeConte, 1861


Pleocoma LeConte, 1856
Pleocoma australis Fall, 1911 (Pleocoma)

Pleocoma badia badia Fall, 1917 (Pleocoma)

Pleocoma badia hirsuta Davis, 1934 (Pleocoma)

Pleocoma nitida Linsley, 1941

Pleocoma venturae Linsley, 1945

Pleocoma behrensii LeConte, 1874 (Pleocoma)

Pleocoma bicolor Linsley, 1935 (Pleocoma)

Pleocoma blaisdelli Linsley, 1938 (Pleocoma)

Pleocoma carinata Linsley, 1938 (Pleocoma)

Pleocoma conjungens conjungens Horn, 1888 (Pleocoma)

Pleocoma conjungens lucia Linsley, 1941 (Pleocoma)

Pleocoma crinita Linsley, 1938 (Pleocoma)

Pleocoma dubitabilis dubitabilis Davis, 1935 (Pleocoma)

Pleocoma dubitabilis leachi Linsley, 1938 (Pleocoma)

Pleocoma fimbriata LeConte, 1856 (Pleocoma)

Pleocoma hirticollis hirticollis Schaufuss, 1870 (Pleocoma)

Pleocoma hirticollis reflexa Hovore, 1972 (Pleocoma)

Pleocoma hirticollis vandykei Linsley, 1938 (Pleocoma)

Pleocoma hoppingi Fall, 1906 (Pleocoma)

Pleocoma hovorei La Rue, 2007 (Pleocoma)

Pleocoma linsleyi Hovore, 1971 (Pleocoma)

Pleocoma marquai Hovore, 1972 (Pleocoma)

Pleocoma minor Linsley, 1938 (Pleocoma)

Pleocoma octopagina Robertson, 1970 (Pleocoma)

Pleocoma oregonensis Leach, 1933 (Pleocoma)

Pleocoma remota Davis, 1934

Pleocoma puncticollis Rivers, 1889 (Pleocoma)

Pleocoma rickseckeri Horn, 1888 (Pleocoma)

Pleocoma rubiginosa rubiginosa Hovore, 1972 (Pleocoma)

Pleocoma rubiginosa transsierrae Hovore, 1977 (Pleocoma)

Pleocoma shastensis Van Dyke, 1933 (Pleocoma)

Pleocoma simi Davis, 1934 (Pleocoma)

Pleocoma sonomae Linsley, 1935 (Pleocoma)

Pleocoma staff Schaufuss, 1870 (Pleocoma)

Pleocoma adjuvans Crotch, 1873

Pleocoma edwardsii LeConte, 1874

Pleocoma ulkei Horn, 1888

Pleocoma trifoliata Linsley, 1938 (Pleocoma)

Pleocoma tularensis Leach, 1933 (Pleocoma)
GEOTRUPIDAE Latreille, 1802

BOLBOCERATINAE Mulsant, 1842

ATHYREINI Howden and Martínez, 1963



Neoathyreus Howden and Martínez, 1963
Neoathyreus fissicornis (Harold, 1880) (Athyreus)

Neoathyreus mixtus Howden, 1964 (Neoathyreus)
BOLBOCERATINI Mulsant, 1842
Bolbelasmus Boucomont, 1911

Kolbeus Boucomont, 1911
Bolbelasmus arcuatus (Bates, 1887) (Bolboceras)

Bolbelasmus bajaensis Howden, 1964 (Bolbelasmus)

Bolbelasmus hornii (Rivers, 1886) (Bradycinetus)

Bolbelasmus minor (Linell, 1896) (Bradycinetus)

Bolbelasmus variabilis Howden, 1964 (Bolbelasmus)
Bolbocerastes Cartwright, 1953
Bolbocerastes imperialis imperialis Cartwright, 1953 (Bolbocerastes)

Bolbocerastes imperialis kansanus Cartwright, 1953 (Bolbocerastes)

Bolbocerastes peninsularis Schaeffer, 1906 (Bradycinetus)

Bolbocerastes regalis Cartwright, 1953 (Bolbocerastes)

Bolbocerastes serratus (LeConte, 1854) (Athyreus)
Bolbocerosoma Schaeffer, 1906
Bolbocerosoma (Bolbocerosoma) Schaeffer, 1906
Bolbocerosoma (Bolbocerosoma) biplagiatum Dawson and McColloch, 1924 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) bruneri Dawson and McColloch, 1924 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) cartwrighti Howden, 1955 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) confusum Brown, 1928 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) elongatum Howden, 1955 (Bolbocerosoma)



Bolbocerosoma (Bolbocerosoma) farctum (Fabricius, 1775) (Scarabaeus)

Scarabaeus cephus Fabricius, 1775

Bolbocerosoma (Bolbocerosoma) hamatum Brown, 1929 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) lepidissimum Brown, 1928 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) mexicanum Howden, 2005 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) pusillum pusillum Dawson and McColloch, 1924 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) pusillum townesi Howden, 1955 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) quadricornum Robinson, 1941 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) ritcheri Howden, 1955 (Bolbocerosoma)

Bolbocerosoma (Bolbocerosoma) tumefactum (Palisot de Beauvois, 1809) (Scarabaeus)
Bolborhombus Cartwright, 1953
Bolborhombus angulus (Robinson, 1947) (Bolboceras)

Bolborhombus magnus Howden, 1964 (Bolborhombus)

Bolborhombus parvulus Cartwright, 1953 (Bolborhombus)

Bolborhombus sallaei (Bates, 1887) (Bolboceras)

Bradycinetus carinatus Schaeffer, 1906

Bolboceras schaefferi Boucomont, 1911
Bradycinetulus Cockerell, 1906

Amechanus Horn, 1870

Bradycinetus Horn, 1871
Bradycinetulus ferrugineus (Palisot de Beauvois, 1809) (Scarabaeus)

Bolboceras furcicollis Laporte, 1840

Bradycinetulus fossatus (Haldeman, 1853) (Bolboceras)

Bradycinetulus rex Cartwright, 1953 (Bradycinetulus)
Eucanthus Westwood, 1848
Eucanthus alutaceus Cartwright, 1944 (Eucanthus)

Eucanthus greeni Robinson, 1948 (Eucanthus)

Eucanthus impressus Howden, 1964 (Eucanthus)

Eucanthus lazarus (Fabricius, 1775) (Scarabaeus)

Scarabaeus meliboeus Fabricius, 1792

Eucanthus mexicanus Howden, 1964 (Eucanthus)

Eucanthus subtropicus Howden, 1955 (Eucanthus)
Odonteus Samouelle, 1819
Odonteus alabamensis Wallis, 1929 (Odonteus)

Odonteus cornigerus (Melsheimer, 1845) (Bolboceras)

Odonteus darlingtoni Wallis, 1928 (Odonteus)

Odonteus falli Wallis, 1928 (Odonteus)

Odonteus filicornis (Say, 1823) (Geotrupes)

Odonteus floridensis Wallis, 1928 (Odonteus)

Odonteus liebecki Wallis, 1928 (Odonteus)

Odonteus obesus LeConte, 1859 (Odonteus)

Odonteus simi Wallis, 1928 (Odonteus)

Odonteus thoracicornis Wallis, 1928 (Odonteus)
GEOTRUPINAE Latreille, 1802
Ceratophyus Fischer von Waldheim, 1823
Ceratophyus gopherinus Cartwright, 1966 (Ceratophyus)
Ceratotrupes Jekel, 1865
Ceratotrupes bolivari Halffter and Martínez, 1962 (Ceratotrupes)
Geotrupes Latreille, 1797
Geotrupes (Anoplotrupes) Jekel, 1865

Melanotrupes Blanchard, 1888

Geotrupes (Cnemotrupes) Jekel, 1865

Onychotrupes Jekel, 1865

Geotrupes (Geohowdenius) Zunino, 1984

Geotrupes (Geotrupes) Latreille, 1797

Geotrupes (Megatrupes) Zunino, 1984
Geotrupes (Anoplotrupes) balyi Jekel, 1865 (Geotrupes)

Geotrupes similis Jekel, 1865

Geotrupes (Cnemotrupes) blackburnii blackburnii (Fabricius, 1781) (Scarabaeus)

Geotrupes conicollis Jekel, 1865

Geotrupes jekelii Horn, 1868

Geotrupes (Cnemotrupes) blackburnii excrementi Say, 1823 (Geotrupes)

Geotrupes (Megatrupes) cavicollis Bates, 1887 (Geotrupes)

Geotrupes (Geohowdenius) egeriei Germar, 1824 (Geotrupes)

Geotrupes lecontei Jekel, 1865

Geotrupes (Anoplotrupes) hornii Blanchard, 1888 (Geotrupes)

Geotrupes (Geohowdenius) opacus Haldeman, 1853 (Geotrupes)

Geotrupes haldemani Jekel, 1865

Geotrupes chevrolati Jekel, 1865

Geotrupes (Cnemotrupes) semiopacus Jekel, 1865 (Geotrupes)

Geotrupes melsheimeri Jekel, 1865

Geotrupes ovalipennis Jekel, 1865

Geotrupes (Cnemotrupes) splendidus miarophagus Say, 1823 (Geotrupes)

Geotrupes (Cnemotrupes) splendidus splendidus (Fabricius, 1775) (Scarabaeus)

Geotrupes gilnickii Jekel, 1865

Geotrupes starki Jekel, 1865

Geotrupes mixtus Horn, 1868

Geotrupes (Geotrupes) stercorarius (Linnaeus, 1758) (Scarabaeus)

Geotrupes (Cnemotrupes) ulkei Blanchard, 1888 (Geotrupes)
Mycotrupes LeConte, 1866
Mycotrupes cartwrighti Olson and Hubbell, 1954 (Mycotrupes)

Mycotrupes gaigei Olson and Hubbell, 1954 (Mycotrupes)

Mycotrupes lethroides (Westwood, 1837) (Geotrupes)

Mycotrupes pedester Howden, 1954 (Mycotrupes)

Mycotrupes retusus (LeConte, 1866) (Geotrupes)

Geotrupes aeneus Felsche, 1909
Peltotrupes Blanchard, 1888
Peltotrupes profundus (Howden, 1952) (Geotrupes)

Geotrupes chalybaeus LeConte, 1878

Peltotrupes dubius Howden, 1955

Peltotrupes youngi Howden, 1955 (Peltotrupes)
PASSALIDAE Leach, 1815

PASSALINAE Leach, 1815

PROCULINI Kaup, 1868
Heliscus Zang, 1906
Heliscus tropicus (Percheron, 1835) (Passalus)
Odontotaenius Kuwert, 1896
Odontotaenius disjunctus (Illiger, 1800) (Passalus)

Passalus cornutus Fabricius, 1801

Passalus distinctus Weber, 1801

Passalus bos Kuwert, 1891

Odontotaenius floridanus Schuster, 1994 (Odontotaenius)

Odontotaenius striatopunctatus (Percheron, 1835) (Passalus)
PASSALINI Leach, 1815
Passalus Fabricius, 1792

Pertinax Kaup, 1869

Neleides Kaup, 1869

Petrejus Kaup, 1869

Mitrorhinus Kaup, 1871

Rhodocanthopus Kaup, 1871

Epiphanus Kaup, 1871

Rhagonocerus Kaup, 1871

Valerius Kuwert, 1891

Cassius Kuwert, 1891

Lucilius Kuwert, 1891

Pertinacides Kuwert, 1891

Epipertinax Kuwert, 1891

Ninoides Kuwert, 1891

Neleurops Kuwert, 1891

Manlius Kuwert, 1891

Phoronaeosomus Kuwert, 1891

Tetraracus Kuwert, 1891

Lophocephalus Kuwert, 1896

Aponelides Kuwert, 1896

Parapertinax Kuwert, 1896

Morosophus Kuwert, 1896

Severus Kuwert, 1896

Polyacanthopus Kuwert, 1896

Synesius Kuwert, 1896

Oeneus Kuwert, 1896

Ptychotrichus Kuwert, 1896

Calidas Kuwert, 1896

Toxeutotaenius Kuwert, 1896

Gnomon Zang, 1905

Lasioperix Zang, 1905

Phaulothorax Zang, 1905

Scalmus Zang, 1905

Psilomus Zang, 1905

Macrolobus Zang, 1905

Alocerus Zang, 1905

Epipleurothrix Zang, 1905

Zosterothrix Zang, 1905

Procululus Zang, 1905

Epiphoroneus Arrow, 1907
Passalus punctatostriatus Percheron, 1835 (Passalus)

Passalus punctiger LePeletier and Serville, 1825 (Passalus)

Passalus punctatissimus Eschscholtz, 1829

Passalus striolatus Eschscholtz, 1829

Neleus laborator Kaup, 1869

Neleus transvalensis Kuwert, 1891

Neleus guatemalae Kuwert, 1891

Neleus difficilis Kuwert, 1891

Neleus distinguendus Kuwert, 1891

Neleus laeviclypeatus Kuwert, 1891

Neleus scurra Kuwert, 1891

Neleus sanio Kuwert, 1891

Neleus chilensis Kuwert, 1891

Neleus nicaraguae Kuwert, 1891

Neleus scelus Kuwert, 1891

Neleus vagans Kuwert, 1891

Neleus scepticus Kuwert, 1891

Neleus sulcicornis Kuwert, 1898

Neleus arcuatotaeniatus Kuwert, 1898

Neleus distinctus Kuwert, 1898

Neleus intermissus Kuwert, 1898

Neleus subcarinaefrons Kuwert, 1898

Neleus subcarinatus Kuwert, 1898

Neleus taeniolatus Kuwert, 1898

Neleus dislocandus Kuwert, 1898

Neleus scutellosulcatus Kuwert, 1898

Neleus durangi Kuwert, 1898

Neleus obtusecornutus Kuwert, 1898

Neleus altidens Kuwert, 1898

Neleus carinaefrons Kuwert, 1898

Neleus acarinatus Kuwert, 1898

Neleus dilatipunctatus Kuwert, 1898

Neleus dispar Kuwert, 1898

Neleus dilatus Kuwert, 1898

Neleus aequatoris Kuwert, 1898

Neleus difformis Kuwert, 1898

Neleus argentinus Kuwert, 1898

Neleus boliviensis Kuwert, 1898

Neleus dilatidentatus Kuwert, 1898

Neleus disjunctus Kuwert, 1898

Neleus approximatidentatus Kuwert, 1898

Neleus dispositus Kuwert, 1898

Neleus labroexcisus Kuwert, 1898

Neleus scurroides Kuwert, 1898

Neleus arrogans Kuwert, 1898

Neleus festae Rosmini, 1902

Neleus cognettii Pangella, 1905

Passalus riograndensis Luederwaldt, 1931
Ptichopus Kaup, 1869
Ptichopus angulatus (Percheron, 1835) (Passalus)

Passalus nodus Apetz, 1837

Passalus thoracicus Smith, 1852

Ptichopus inca Kuwert, 1891

Ptichopus nitidus Kuwert, 1891

Ptichopus montezuma Kuwert, 1891

Ptichopus aberrator Kuwert, 1891

Ptichopus inflatus Kuwert, 1898

Ptichopus borellii Rosmini, 1902
TROGIDAE MacLeay, 1819
Omorgus Erichson, 1847

Chesas Burmeister, 1876

Lagopelas Burmeister, 1876

Megalotrox De Borre, 1886
Omorgus asper LeConte, 1854 (Omorgus)

Omorgus carinatus (Loomis, 1922) (Trox)

Omorgus fuliginosus (Robinson, 1941) (Trox)

Omorgus howelli (Howden and Vaurie, 1957) (Trox)

Omorgus inflatus (Loomis, 1922) (Trox)

Omorgus monachus (Herbst, 1790) (Trox)

Trox tuberculatus Palisot de Beauvois, 1818

Omorgus pustulatus LeConte, 1854

Omorgus nodosus (Robinson, 1940) (Trox)

Omorgus punctatus (Germar, 1824) (Trox)

Omorgus integer LeConte, 1854

Omorgus morsus LeConte, 1854

Omorgus rubricans (Robinson, 1946) (Trox)

Omorgus scabrosus (Palisot de Beauvois, 1818) (Trox)

Omorgus scutellaris (Say, 1824) (Trox)

Omorgus suberosus (Fabricius, 1775) (Trox)

Trox crenatus Olivier, 1789

Trox gibbus Olivier, 1789

Trox ovatus Palisot de Beauvois, 1818

Trox denticulatus Palisot de Beauvois, 1818

Trox alternatus Say, 1831

Trox nobilis Wollaston, 1867

Trox tricolor Blackburn, 1904

Trox manilensis Schultze, 1915

Trox torressalai Baguena, 1959

Trox triestinae Pittino, 1987

Omorgus tesselatus LeConte, 1854 (Omorgus)

Trox nigrescens Robinson, 1941

Omorgus texanus LeConte, 1854 (Omorgus)

Omorgus suturalis LeConte, 1854

Trox oligonus Loomis, 1922

Omorgus tomentosus (Robinson, 1941) (Trox)

Omorgus tytus (Robinson, 1941) (Trox)

Omorgus umbonatus LeConte, 1854 (Omorgus)

Trox platycyphus Loomis, 1922

Trox confluentus Robinson, 1941
Trox Fabricius, 1775

Pseudotrox Robinson, 1948
Trox aequalis Say, 1832 (Trox)

Trox affinis Robinson, 1940 (Trox)

Trox atrox LeConte, 1854 (Trox)

Trox capillaris Say, 1824 (Trox)

Trox contractus Robinson, 1940 (Trox)

Trox fascifer LeConte, 1854 (Trox)

Trox floridanus Howden and Vaurie, 1957 (Trox)

Trox foveicollis Harold, 1872 (Trox)

Trox aciculeatus Robinson, 1940

Trox frontera Vaurie, 1955 (Trox)

Trox gemmulatus Horn, 1874 (Trox)

Trox hamatus Robinson, 1940 (Trox)

Trox laticollis LeConte, 1854 (Trox)

Trox plicatus Robinson, 1940 (Trox)

Trox robinsoni Vaurie, 1955 (Trox)

Trox scaber (Linnaeus, 1767) (Silpha)

Trox barbosus von Laicharting, 1781

Trox arenarius Fabricius, 1787

Trox arenosus Gmelin, 1790

Trox hispidus von Paykull, 1798

Trox trisulcatus Curtis, 1844

Trox niponensis Lewis, 1895

Trox sonorae LeConte, 1854 (Trox)

Trox alternans LeConte, 1854

Trox lecontei Harold, 1872

Trox sordidus LeConte, 1854 (Trox)

Trox spinulosus dentibius Robinson, 1940 (Trox)

Trox spinulosus spinulosus Robinson, 1940 (Trox)

Trox spinulosus simi Robinson, 1940 (Trox)

Trox striatus Melsheimer, 1845 (Trox)

Trox terrestris Say, 1825 (Trox)

Trox tuberculatus (De Geer, 1774) (Scarabaeus)

Trox serrulatus Palisot de Beauvois, 1818

Trox canaliculatus Say, 1824

Trox unistriatus Palisot de Beauvois, 1818 (Trox)

Trox porcatus Say, 1825

Trox variolatus Melsheimer, 1845 (Trox)

Trox erinaceus LeConte, 1854

Trox monticola Robinson, 1940
GLARESIDAE Kolbe, 1905
Glaresis Erichson, 1848

Eoglaresis Semenov and Medvedev, 1932

Eremoglaresis Semenov and Medvedev, 1932

Afroglaresis Petrovitz, 1968
Glaresis arenata Gordon, 1974 (Glaresis)

Glaresis canadensis Brown, 1928 (Glaresis)

Glaresis cartwrighti Gordon, 1970 (Glaresis)

Glaresis clypeata Van Dyke, 1928 (Glaresis)

Glaresis confusa Brown, 1928 (Glaresis)

Glaresis dakotensis Gordon, 1970 (Glaresis)

Glaresis ecostata Fall, 1907 (Glaresis)

Glaresis gordoni Warner, 1995 (Glaresis)

Glaresis howdeni Gordon, 1970 (Glaresis)

Glaresis inducta Horn, 1885 (Glaresis)

Glaresis knausi Brown, 1928

Glaresis medialis Gordon, 1969 (Glaresis)

Glaresis mendica Horn, 1885 (Glaresis)

Glaresis phoenicis Fall, 1907 (Glaresis)

Glaresis texana Gordon, 1970 (Glaresis)

Glaresis zvirgzdinsi Warner, 1995 (Glaresis)
DIPHYLLOSTOMATIDAE Holloway, 1972
Diphyllostoma Fall, 1901

Phyllostoma Fall, 1901
Diphyllostoma fimbriatum (Fall, 1901) (Phyllostoma)

Diphyllostoma linsleyi Fall, 1932 (Diphyllostoma)

Diphyllostoma nigricollis Fall, 1912 (Diphyllostoma)
LUCANIDAE Latreille, 1804

AESALINAE MacLeay, 1819

NICAGINI LeConte, 1861
Nicagus LeConte, 1861
Nicagus occultus Paulsen and Smith, 2005 (Nicagus)

Nicagus obscurus (LeConte, 1847) (Ochodaeus)
LUCANINAE Latreille, 1804

LUCANINI Latreille, 1804
Dorcus MacLeay, 1819
Dorcus brevis (Say, 1825) (Lucanus)

Dorcus caucasicus Ganglbauer, 1886

Dorcus parallelus (Say, 1824) (Lucanus)

Lucanus oblongus de Charpentier, 1825

Dorcus costatus LeConte, 1866

Dorcus nanus Casey, 1909

Dorcus carnochani Angell, 1916
Lucanus Scopoli, 1763

Hexaphyllus Mulsant, 1839

Pseudolucanus Hope, 1845
Lucanus capreolus (Linnaeus, 1764) (Scarabaeus)

Lucanus dama Fabricius, 1775

Lucanus muticus Thunberg, 1806

Lucanus trigonus Thunberg, 1806

Lucanus elaphus Fabricius, 1775 (Lucanus)

Lucanus carlengi Angell, 1916

Lucanus mazama (LeConte, 1861) (Dorcus)

Pseudolucanus bicostatus Angell, 1917

Lucanus placidus Say, 1825 (Lucanus)

Lucanus lentus Laporte, 1840
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