How to Know Pennsylvania’s Biota Odonata

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How to Know Pennsylvania’s Biota
Odonata (Dragonflies and damselflies) of Pennsylvania
There is no formal publication dealing solely with PA.
Dunkle, Sidney W.  2000 Dragonflies through Binoculars - A Field Guide to the Dragonflies of North America. Oxford University Press, 266 p. ISBN 0-19-511268-7.  $19.97 (Amazon).
Glotzhober, R.C.  and D. McShaffrey (Editors).  2002.  The Dragonflies and Damselflies of Ohio.  Ohio Biological Survey Bulleting New Series Volume 14 Number 2.  ix + 364 p. ISBN 0-86727-145-0. $40.

Includes generic keys to the larvae and specific keys to the adult males and females for all 162 species known from Ohio (as well as a few species likely to occur there).

Lam, Ed. Damselflies of the Northeast. Biodiversity Books, PO Box 353, Eastchester, NY 10709. ISBN 0-9754015-0-5. $20.
Nikula, B., J. Loose, and M. Burne. 2003. A Field Guide to the Dragonflies and Damselflies of Massachusetts. Massachusetts Natural Heritage Program, 1 Rabbit Hill Road, Westborough, MA 01581. 197 pp. 20.
Nikula, B. and J. Sones. 2002. Stokes Beginners Guide to Dragonflies and Damselflies. Little, Brown & Company, Boston. 160 pp. ISBN 0-316-81679-5. $8.95
Rosche, L. 2002. Dragonflies and Damselflies of Northeast Ohio. Cleveland Museum of Natural History. Cleveland, OH. 94 pp. ISBN 0-9717460-0-1. $18.95.


Bob Moul:

Tom Pawlesh

Checklist of PA Odonates:

Odonates of Pennsylvania

Ohio Odonata survey.

Beatty, G.H. and A.F. Beatty. 1968. Checklist and bibliography of Pennsylvania Odonata. Pennsylvania Academy of Science 42: 120-129.

Beatty, G.H. and A.F. Beatty. 1971a. The distribution of Pennsylvania Odonata. Pennsylvania Academy of Science 45: 147-167.

Beatty, G.H. and A.F. Beatty. 1971b. The Odonata of Pennsylvania: List and cross-references to literature containing data on occurrence in the state. Pennsylvania Academy of Science 45: 111- 120.

Beatty, G.H. and A.F. Beatty and H.B. White III. 1969. Seasonal distribution of Pennsylvania Odonata. Pennsylvania Academy of Science 43: 119-123.

Donnelly, T.W. 1992. The Odonata of New York State. Bulletin of American Odonatology 1(1): 1-27.

Lepidoptera of Pennsylvania
There is no formal publication dealing solely with PA.

Allen, T. J. Brock, and J. Glassberg. 2005. Caterpillars in the Field and Garden. Oxford University Press. NY. 232 pp. ISBN 0-19-514987-4. $19.97 (Amazon)

Wagner, D. L. 2005. Caterpillars of Eastern North America. Princeton University Press. Princeton, NJ. ISBN 0-691-12144-3. 512 pp. $19.77 (Amazon)
Wagner, D. L., V. Giles, R. Reardon, and M. McManus. Caterpillars of Eastern Forests. 1997. Forest Health Technology Enterprise Team. Morgantown, WV. FHTET-96-34.
Wright, A. B. 1993. Peterson First Field Guide to Caterpillars. Peterson First Guides. Houghton-Mifflin. Boston. 128 pp. ISBN 0-395-56499-9. $5.95
Allen, T. 1997. The Butterflies of West Virginia and Their Caterpillars. University of Pittsburgh Press.. 388 pp ISBN 0-8229-5657-8. $18.96 (Amazon)
Brock, J. and K. Kaufman. 2003. Butterflies of North America. Houghton Mifflin. 384 pp. ISBN 0-618-15312-8. $14.96 (Amazon)
Glassberg, J. 1999. Butterflies Through Binoculars: The East. Oxford University Press. NY. 242 pp. ISBN 0-19-510668-7.
Iftner, D., J. Shuey, and J. Calhoun. 1992. Butterflies and Skippers of Ohio. Ohio Biol. Surv. Bull. New Seris Vol. 9 No. 1, xii +212 pp. ISBN 0867271078
Mitchell, R. and H. Zim. 2001 ed. Butterflies and Moths. 160 pp. Golden Guides from St. Martin's Press. ISBN 1582381364. $6.95.
Opler, P. and V. Malikul. 1992. Eastern Butterflies. Peterson Field Guide Series. Houghton-Mifflin. ISBN 0-395-63279-X. 396 pp. $16.95.
Wright, D. 2006. Atlas of Pennsylvania Butterflies, 5th ed. Unpublished MS. Available from
Covell, C. 2005. A Field Guide to the Moths of Eastern North America, 2nd ed. Virginia Museum of Natural History. ISBN 1884549217. 518 pp. $40.
Holland, W. 1968. The Moth Book. A guide to the moths of North America, New Ed. Edition. Dover Publishing. 479 pp. ISBN 0486219488
Digital Guide to Moth Identification.

Bob Patterson's Entomology Page,

List of Butterflies of Pennsylvania,

Butterflies and Moths of North America,

Wildflowers of Pennsylvania
Clemants, S. and C. Gracie. 2006. Wildflowers in the Field and Forest. A field guide to the northeastern United States. Princeton University Press. ISBN: 0195150058. 480 pp. $23.10 (Amazon)
Gleason, H. and A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada, 2nd ed. ISBN: 0893273651. 910 pp. $69.
Haywood, M. J. and P. Monk. 2001. Wildflowers of Pennsylvania. Venture Graphics. ISBN 0-9710614-0-8. 407 pp.
Hill, R. 1986. Poisonous Plants of Pennsylvania. Commonwealth of Pennsylvania. ISBN: 0818200782. 175 pp.

Medve, R. and M. Medve. 1990. Edible Wild Plants of Pennsylvania and Neighboring States. Pennsylvania State University Press. ISBN: 0271006978. 256 pp

Newcomb, L. 1989. Newcomb’s Wildflower Guide, Reprint Ed. Little Brown & Co. ISBN: 0316604429. 490 pp. $13.59 (Amazon)
Rhoads, A., T. Block, and A. Anisko. 2000. The Plants of Pennsylvania: An Illustrated Manual. University of Pennsylvania Press. ISBN 0812235355. 1061 pp. $75.
Stokes, D. and L. Stokes. 1986. A Guide to Enjoying Wildflowers, Reprint Ed. Little Brown & Co. ISBN: 0316817317. 371 pp.
Wildflowers of Western Pennsylvania,
Fishes of Pennsylvania
Cooper, Edwin. 1983. Fishes of Pennsylvania and the Northeastern United States. Pennsylvania State Univ. Press. University Park, PA. ISBN: 0271003375. 273 pp. Out of print. Used is selling for $175+.
Denoncourt, R. F. 1975. Key to the families and genera of Pennsylvania freshwater fishes and the species of freshwater fishes of the Susquehanna River drainage above Conowingo Dam. Proc. Pa. Acad. Sci. 49:82.88.
Gallery of Pennsylvania Fishes. Text and photos from Pennsylvania Fishes.
Steiner, L. Pennsylvania Fishes. Pennsylvania Fish & Boat Commission.

Aquatic Sciences
Voshell, J. 2002. A Guide to Common Freshwater Invertebrates of North America. McDonald and Woodward Publishing Company ISBN: 0939923874. 456 pp.

Birds of Pennsylvania

There may be a publication for the county where you live. Check here:

Or here

Brauning, D., ed. 1992. Atlas of Breeding Birds in Pennsylvania. University of Pittsburgh Press ISBN 0822936925. 484 pp. Out of print, but available online at

Haas, F. and B. Haas. 2005. Annotated List of the Birds of Pennsylvania, 2nd Edition. Pennsylvania Society for Ornithology. Available from the society’s website, $7.00.

McWilliams, G. and D. Brauning. 1999. The Birds of Pennsylvania. Cornell University Press. Ithaca, NY. ISBN 0-8014-3643-5. 479 pp. $45.
Pennsylvania Society for Ornithology

Official PA bird list-

Fergus, C. 2000. Wildlife of Pennsylvania and the Northeast. Stackpole Books. Mechanicsburg, PA. ISBN 0-8117-2899-4. 438 pp. $19.95.

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