Drafts: V. N. Gladkova [VG], B. A. Yurtsev [BY], V. N. Tikhomirov [VT] and R. Elven [RE]




Дата канвертавання22.04.2016
Памер52.55 Kb.
59  Rosaceae (Drafts: V.N. Gladkova [VG], B.A. Yurtsev [BY], V.N. Tikhomirov [VT] and R. Elven [RE])

S Spiraeaceae; Malaceae; Amygdalaceae

Comments:

(1) We have not decided on any order of genera in Rosaceae. I have tentatively applied with some modifications the traditional order of Flora Europaea with four subfamilies   Spiraeoideae, Rosoideae, Maloideae and Prunoideae   even if these are not to be formally entered in the draft. (Elven)



59.1  Spiraea L. (1753), Sp. Pl. 489. [VG]

Comments:

(1) An additional species is indicated by Gladkova in Yurtsev (1984, Fl. Arct. URSS 9, 1) from lower Lena in N Siberia but probably not in the Arctic as we delimit it: S. dahurica (Rupr.) Maxim. (1879), Trudy Glavn. Bot. Sada 6: 190; basionym: S. alpina Pall. var. dahurica Rupr. (1857), Bull. Phys.-Math. Acad. Sci. (Petersb.) 15: 362. Please state whether it should be included. (Elven)
59.1.1  Spiraea salicifolia L. (1753), Sp. Pl. 489.

S

2n= 36 (4x).



2nD Sax (1936).

G SIB RFE

Comments:

(1) In addition to arctic occurrence in Kharaulakh and W Chukotka this species also approaches the Arctic in S Chukotka. (Gladkova)


59.1.2  Spiraea beauverdiana Schneid. (1905), Bull. Herb. Boiss., sér. 2, 5: 348.

S ?S. beauverdiana Schneid. var. stevenii Schneid. (1905), Bull. Herb. Boiss., sér. 2, 5: 348; ?S. stevenii (Schneid.) Rydb. (1908), N. Amer. Fl. 22: 247.

T [Described from N Japan: Riishiri I.]

2n= (1) 14 (2x). (2) 18 (2x).

2nD (1) ? (2) Sokolovskaya (1960a); Zhukova (1967a); Zhukova & Tikhonova (1973). The two latter are cited by Löve & Löve for S. stevenii.

G RFE ALA CAN

Comments:

(1) The question is whether we have two races   an Asian ('beauverdiana') and a North American ('stevenii')   or only minor variation. The 'stevenii' entity has been described as species and variety. If the two are 'continental' races, subspecies is the obvious choice. However, var. stevenii may only have been described to encompass small plants in northern or mountainous areas. Spiraea stevenii is given for Russian Far East by Löve & Löve (1975) and included in Gladkova's draft from Beringian Alaska, North Alaska and Central Canada. Rydberg used this name for all the American plants and by him it seems to be a simple North American synonym for S. beauverdiana. (Elven)



WARNING! Perhaps two races (subspecies).
59.1.3  Spiraea media F. Schmidt (1792), Oesterr. Allgem. Baumzeit. 1: 53.

S

2n= (1) 10. (2) 18 (2x).



2nD (1) Baksay (1957b C Eur). (2) ?

G RUS SIB

Comments:

59.2  Luetkea Bong. (1832), Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 130, t. 2. [RE]
59.2.1  Luetkea pectinata (Pursh) Kuntze (1891), Revis. Gen. Pl. 1: 217.

B Saxifraga pectinata Pursh (1814), Fl. Amer. Sept. 1: 312.

S

T [Described from: "On the north-west coast", i.e. Alaska.]



2n=

2nD


G ALA? CAN?

Comments:

(1) Genus and species added to the Russian drafts. A border case in SW Alaska (Kuskokwin Mts.) and the lowermost Mackenzie River valley, see Hultén (1968) and Porsild & Cody (1980).

59.3  Aruncus L. (1758), Opera Var. 259. [VG]
59.3.1  Aruncus dioicus (Walter) Fern. (1939), Rhodora 41: 423.

B Actaea dioica Walter (1788), Fl. Carol. 152.

S ?A. sylvester Kostel. (1844), Ind. Hort. Bot. Prag. 15; ?A. kamtschaticus (Maxim.) Rydb. (1908), N. Amer. Fl. 22: 256 [basionym: A. sylvester Kostel. var. (gamma) kamtschaticus Maxim. (1879), Trudy Glavn. Bot. Sada 6: 170].

2n= 18 (2x).

2nD Löve & Löve (1975) list several counts, two Russian as arctic.

G SIB RFE

Comments:

(1) Gladkova proposes A. kamtschaticus. The global phytogeographical pattern of this species or species-group is repeated in many other genera. It occurs disjunct in major mountainous areas in Europe, Asia and N America, in the present case as a series from S/C Europe through Caucasus, S/C Asia, China, NE Asia, and ending in NW America. The cases differ, of course, in degrees of differentiation. The same do the treatments. North Americans oten tend to accept one widely delimited species, the Russians aggregates of related species, and Hultén and other W Europeans subspecies. Look also, e.g., at the treatments of the Astragalus australis and Anemone narcissiflora complexes.

In this case, the basic Russian treatment is Pojarkova's in Fl. SSSR. She restricted A. sylvester Kostel to the Caucasus populations, described the plants of S/C Asia, China and southern Far East as A. asiaticus and accepted the Maximovicz   Rydberg name A. kamtschaticus for the NE Asiatic plants. Rydberg (1908) himself regarded the North American plants as a A. acuminatus Rydb. Hultén (1946) discussed the case and found difficult to accept specific entities based on the variable criteria indicated.

It might therefore be better to include this plant in a more widely defined A. dioicus with a comment. Subspecific treatment may be preferable but I have not found relevant names to propose at that level. (Elven)



WARNING! Subspecies or a separate NE Asiatic species might be recognised.

59.4  Filipendula Mill. (1754), Gard. Dict. Abr., ed. 4, no. 1. [VG]
59.4.1  Filipendula ulmaria (L.) Maxim. (1879), Trudy Glavn. Bot. Sada 6: 251.

B Spiraea ulmaria L. (1753), Sp. Pl. 490.

Comments:

(1) Two varieties are tentatively included. Gladkova's draft includes var. denudata, and for formal reasons the rest then becomes var. ulmaria. Var. denudata could perhaps be reduced to a comment? It seems to be very weakly characterised morphologically and does not seem to have the distributional characteristic needed of a subspecies. (Elven)



WARNING! The varieties might be omitted.
59.4.1.1  Filipendula ulmaria (L.) Maxim. var. ulmaria

S

2n= 14 (2x).



2nD Löve & Löve (1975) list several non-arctic counts.

G ICE NOR RUS SIB?

Comments:

(1) Recorded in the vicinities of ES.5 (sw) [Taimyr]. (Gladkova)


59.4.1.2  Filipendula ulmaria (L.) Maxim. var. denudata (J. Presl & C. Presl) Beck ***

B Spiraea denudata J. Presl & C. Presl (1819), Fl. Cech. 101.

S F. denudata (J. Presl & C. Presl) Fritsch (1889), Verh. Zool.-Bot. Ges. Wien 39: 591; F. ulmaria (L.) Maxim. subsp. denudata (J. Presl & C. Presl) Hayek ***

2n=


2nD

G RUS


Comments:

(1) Intermediate forms between var. denudata and var. ulmaria are not rarely found, especially in East Siberia. (Gladkova)



59.5  Rubus L. (1753), Sp. Pl. 492. [VG]
59.5.1  Rubus chamaemorus L. (1753), Sp. Pl. 494.

S

2n= 56 (8x).



2nD Löve & Löve (1975) list numerous counts, four as arctic.

G NOR RUS SIB RFE ALA CAN GRL

Comments:
59.5.2  Rubus arcticus L. (1753), Sp. Pl. 494.

S

Comments:



(1) Gladkova treats all the entities 'arcticus', 'acaulis', 'stellatus' and 'decipiens' as full species in her draft. Species rank is also accepted by Porsild & Cody (1980) and some other North American sources. Hultén (1946) did the same but later he changed his opinion. Hultén (1968) and Hultén & Fries (1986) apply subspecies for the three former entities and argue that hybrid swarms are formed where they overlap. The last entity   'decipiens' - is such a hybrid swarm in his concept.

If the three former are overlapping and intergrading parapatric entities, rank of subspecies is more appropriate that species rank according to our general rules. At the same time there are evident morphological differences among the extremes. My tentative proposal here is subspecies. If we decide to treat them as species, we must have a R. arcticus aggregate with a comment on the intermediates. The 'decipiens' entity has been reduced to a comment as there also can be doubts about its name. (Elven)



WARNING! The entities here entered as subspecies could also be considered as species.
59.5.2.1  Rubus arcticus L. subsp. arcticus

S

2n= (1) 14 (2x). (2) 21 (3x).



2nD (1) Löve & Löve (1975) list several counts, four as arctic. (2) ?

G NOR RUS SIB RFE ALA CAN

Comments:

(1) Var. [beta] grandiflorus Ledeb. (1844), Fl. Ross. 2: 70   CW (f), CS (f), CE (f), AW (s?)   a few records are also known in Siberia (the distribution is to be checked). (Gladkova)

(2) Due to uncertainties, I have not yet included this variety as a formal entity. (Elven)
59.5.2.2  Rubus arcticus L. subsp. acaulis (Michx.) Focke ***

B R. acaulis Michx. (1803), Fl. Bor.-Amer. 1: 298.

S R. arcticus L. subsp. stellatus (Sm.) Boivin emend. Hultén var. acaulis (Michx.) Boivin (1955), Bull. Soc. Bot. France 102: 235; R. arcticus L. var. grandiflorus auct. amer., non Ledeb. (1844).

T [Described from Canada: the Hudson Bay area.]

2n= 14 (2x).

2nD Löve & Ritchie (1966); Larsson (1969).

G ALA CAN

Comments:

(1) This American entity is also tentatively indicated for Chukotka by Hultén (1968) but not confirmed. (Elven)
59.5.2.3  Rubus arcticus L. subsp. stellatus (Sm.) Boivin (1955), Bull. Soc. Bot. France 102: 235 [quoad var. stellatus?] emend. Hultén ***

B R. stellatus Sm. (1791), Pl. Icon. Ined. 3: t. 64.

S

T [Described from S Alaska, "Foggy Harbour".]



2n= 14 (2x).

2nD Vaarama (1954); Larsson (1969).

G RFE ALA

Comments:

(1) Rubus decipiens (Gladkova) Gladkova *** [not in Index kewensis], basionym: R. stellatus Sm. var. decipiens Gladkova (1980), Novosti Sist. Vyssh. Rast. 17: 155, is proposed as an additional species in Gladkova's draft. The name seems to be a later homonym for R. decipiens P.J. Muell. (1859), Pollichia 16-17: 158, and therefore inapplicable at level of species. It is interpreted as a possible hybrid between subsp. arcticus and subsp. stellaris (in Gladkova's draft R. arcticus s. str. x R. stellaris) and is probably best treated as a hybrid without a specific entry. It should be commented on as it is included, e.g., in Fl. Arct. URSS. We have at least two similar hybrid complexes that should be treated in the same way, see below: R. x paracaulis (R. arcticus subsp. acaulis x R. pubescens) and R. x castoreus (R. arcticus subsp. arcticus x R. saxatilis). (Elven)
The Rubus saxatilis aggregate (R. pubescens, R. saxatilis)

Comments:

(1) Two closely related but fully allopatric entities traditionally treated as species. Subspecies might be an alternative. They also have the same hybridisation pattern with their respective counterparts in the R. arcticus group. (Elven)

WARNING! The two species might be reduced to subspecies.
59.5.3  Rubus saxatilis L. (1753), Sp. Pl. 494.

S

2n= 28 (4x).



2nD Löve & Löve (1975) list numerous counts, three as arctic.

G ICE NOR RUS GRL

Comments:

(1) Hybrids between R. arcticus subsp. arcticus and R. saxatilis may form large, persistent stands and sometimes occur outside the range of the former parent. They have been named as R. castoreus Laest. (1839), Nov. Acta Upsal. 11: 246, 296, but should be treated as primary hybrids with clonal spread and not as a separate taxon. (Elven)


59.5.4  Rubus pubescens Raf. (1811), Med. Repos. New York, ser. 3, 2: 333.

S R. saxatilis L. var. canadensis Michx. ***

T [Described from Canada: the Hudson Bay area.]

2n=


2nD

G CAN


Comments:

(1) Also in the vicinity of 'Baffin-Labrador' in the southern Labrador Peninsula. (Gladkova)

(2) Gladkova accepts Rubus paracaulis L.H. Bailey (1941), Gentes Herbarium 5: 33, basionym: R. pubescens Raf. var. paracaulis (Bailey) Boivin ***, as a species. It is interpreted as the hybrid between R. arcticus subsp. acaulis and R. pubescens and has a scattered occurrence in the large area of overlap between the proposed parents in Canada. It seems to be a very close parallel to the European R. castoreus and should be treated in the same way, as a hybrid without formal entry but with a comment. (Elven)
59.5.5  Rubus humulifolius C.A. Mey. (1848), Beitr. Pflanzenk. Russ. Reiches 5: 57.

S

2n= 14 (2x).



2nD ***

G RUS SIB

Comments:
59.5.6  Rubus idaeus L. (1753), Sp. Pl. 492.

S

Comments:



(1) Two or three subspecies? Subsp. sachalinensis is united with subsp. melanolasius by some authors, e.g., by Hultén (1946) as he extends it east to NE America. (Elven)
59.5.6.1  Rubus idaeus L. subsp. idaeus

S

2n= 14 (2x).



2nD Löve & Löve (1975) list numerous non-arctic counts; Lövkvist & Hultgård (1999 Sweden).

G NOR RUS GRL*

Comments:
59.5.6.2  Rubus idaeus L. subsp. sachalinensis (Lévl.) Focke (1911), Spec. Ruborum 2: 210.

B R. sachalinensis Lévl. (1909), Feddes Repert. 6: 352 [p. 332 acc. to Hultén 1946].

S

T [Described from Sakhalin.]



2n=

2nD


G RUS SIB

Comments:


59.5.6.3  Rubus idaeus L. subsp. melanolasius (Dieck.) Focke (1896), Abh. Naturwiss. Ver. Bremen 13, 3: 472.

B Rubus melanolasius Dieck. ***

S ?R. strigous Michx. (1803), Fl. Bor.-Amer. 1: 297.

2n= 14 (2x).

2nD Löve & Löve (1975) list four non-arctic counts.

G ALA CAN

Comments:

(1) Index Kewensis has another author citation for the subspecies and also for the species basionym. (Elven)

(1) A border case in W Alaska, in the Yukon River delta (Hultén 1968). Mapped into N Alaska (Brooks Range) by Hultén (1968). These two regions are added to Gladkova's draft. (Elven)

59.6  Rosa L. (1753), Sp. Pl. 491. [VG]

Comments:

(1) Hultén (1968) maps R. woodsii Lindl. (1820), Ros. Monogr. 21, very close to the Arctic in three places near S Hudson Bay and James Bay. Scoggan (1978) seems, however, only to list sites south of the Arctic. Not included yet. (Elven)
59.6.1  Rosa acicularis Lindl. (1820), Ros. Monogr. 44, t. 8.

S

T [Described from Siberia.]



Comments:

(1) Hultén & Fries (1986): subsp. sayi in RFE ALA CAN. Recognized as species by Löve & Löve (1975) and in Gladkova's draft. Gladkova also draws the limit between 'acicularis' and 'sayi' more to the east, including only North American plants in 'sayi'. This may indicate some problems in separation and be a justification for subspecies rather than species. (Elven)


59.6.1.1  Rosa acicularis Lindl. subsp. acicularis

S

2n= (1) 42 (6x). (2) 56 (8x).



2nD (1) ? (2) Löve & Löve (1975) list numerous non-arctic counts.

G RUS SIB RFE ALA

Comments:
59.6.1.2  Rosa acicularis Lindl. subsp. sayi (Schwein.) Lewis (1959), Brittonia 11: 19 [***CHECK THIS!***]

B R. sayi Schwein. in Long. (***), Exped. Winnep. 2: 388.

S R. acicularis Lindl. var. sayiana Erlans. (1926), Papers Mich. Acad. Sci. Arts Lett. 5: 85.

2n= 42 (6x).

2nD Löve & Löve (1975) list several non-arctic counts.

G ALA CAN

Comments:
59.6.2  Rosa majalis Herrm. (1762), Diss. Rosa 8.

S R. cinnamomea L. (1759), non L. (1753).

2n= 14 (2x).

2nD Löve & Löve (1975) list numerous non-arctic counts.



G RUS

Comments:


База данных защищена авторским правом ©shkola.of.by 2016
звярнуцца да адміністрацыі

    Галоўная старонка