Draba taimyrensis




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53.23.17  Draba taimyrensis Tolm. (1975), Fl. Arct. URSS 7: 135.

S

T Siberia: Ad sinum Jamu Baikura lacu Taimyrensi, 12.08.1947, M.I.Wellikainen (LE), holotype.



2n=

2nD


G SIB

Comments:

(1) Known only from a small area in Taimyr Peninsula. The material available (LE) is fairly distinct but at the same time there is much variation in D. 'lactea/wahlenbergii'. Validity of the entity as a separate species should therefore be verified. (Elven)

WARNING! Should be more thoroughly studied and justified before acceptance.
?53.23.18  Draba wahlenbergii Hartm. (1820), Handb. Skand. Fl., ed. 1, 249.

S D. pseudopilosa Pohle (1914), Bull. Jard. Bot. Pétersb. 14, 4 6: 469; D. boecheri Gjaerev. & Ryvarden (1977), K. Norske Vidensk. Selsk. Skr. 1977-4: 23; D. lactea auctt., non Adams (1817).

2n= (1) 24 (3x). (2) 36 [or 32?] (4x). (3) 48 (6x).

2nD (1) Böcher in Gjærevoll & Ryvarden (1977 Grl); see comment. (2) Zhukova & Petrovsky (1984 NE As, as D. pseudopilosa); see comment. (3) Jørgensen et al. (1958 Grl); Böcher (1966a Grl & Sb); Knaben & Engelskjøn (1967 Norw); Zhukova (1968 Sib); Zhukova & Tikhonova (1971, 1973 Sib); Zhukova et al. (1973 Sib); Mulligan (1974 N Am); Packer & McPherson (1974 Ala); Zhukova & Petrovsky (1984 NE As, as D. lactea); Zhukova & Petrovsky (1984 NE As, as D. pseudopilosa); Brochmann et al. (1993 Sb)

G NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) The traditional name D. lactea for this species must be rejected for two very good reasons. The description fits D. fladnizensis and explisitely states a different set of characters from that found in the plant currently named D. lactea. The well-preserved specimen available for typification in LE belongs to D. fladnizensis in current usage; the same seems to be the case with the only other possible type specimen (in Moscow).

There is a major arctic species, mainly or exclusively hexaploid, that differs in numerous characters from the diploid D. fladnizensis. The next available name for it is probably D. wahlenbergii Hartm. (1820). This name was explicitely coined to replace D. androsacea sensu Wahlenb. (1812), non Willd (1800). Draba androsacea Willd. is a synonym of what currently is known as Arabis scopoliana Boiss. (1842), also described two other times as a Draba, as D. ciliata Scop. (1772) and as D. ciliaris Host (1797). The next alternative is D. lapponica Willd. ex DC. (1821), Syst. Nat. 2: 344, which was coined for the same reason and also based on the Wahlenberg plant. The three names D. androsacea sensu Wahlenberg (1812), D. wahlenbergii Hartm. (1820) and D. lapponica Willd. ex DC. (1821) should therefore be homotypic. The remaining question is whether there is a Wahlenberg material available for unambiguous typification. We will check in UPS, but until further the Hartman name is introduced here. (Elven & Petrovsky)

(2) A corollary is that North American authors probably have read Adams' description better than the Russians and especially than the N Europeans (e.g., Knaben 1966). This might be a main reason for the traditional North American problems in separation between D. fladnizensis and D. lactea Adams; they are the same. However, Hultén's statement of delimitation problems and frequent intermediates (Hultén 1945, 1968) and Scoggan's (1978) merge of them have no foundation. Natural hybrids have never been proved and artificial hybrids are sterile (Brochmann et al. 1993). (Elven)

(3) The single tetraploid chromosome number should be checked as the tetraploid level is unknown elsewhere in D. 'lactea/wahlenbergii' but known from related species. For the triploid count, see the D. boecheri comment below. (Elven)

(4) An alloploid origin involving two or more diploids of the Series Pilosae, Lacteae and/or Nivales has been proposed several times (see Brochmann et al. 1992b). It seems probable that either D. fladnizensis or D. subcapitata   or both   or their progenitors have been involved. It is now much more dubious whether any species of the D. nivalis aggregate has been involved (Scheen et al. in prep.). The species may well be polyphyletic and should be allowed to include a significant morphological variation. (Elven)

(5) Draba pseudopilosa Pohle is reduced to synonymy. The type (in LE, from the Lena River estuary) falls well within the current concept of this species. (Petrovsky & Elven)

(6) Draba boecheri Gjaerev. & Ryvarden is reduced to synonymy. It was described as a very local SW Greenland endemic and is typified from Greenland: Jensens Nunatakker, Hornblenderyggen (nordsiden), 31.07. 07.08.1967, leg O. Gjærevoll & L. Ryvarden (TRH) holotype. The spelling of the epithet has been changed here from 'böcheri' as given in original publication and on the type sheet. The type specimens deviate in some characters from the majority of D. 'wahlenbergii' but not enough to merit rank as a separate entity. The two other sheets available (all that ever has been collected) partly contain similar plants, partly plants of either D. norvegica Gunn. or D. arctogena (E. Ekman) E. Ekman (both of series Rupestres). The description is conbined from both elements and does not fit either of them very well.

The reported triploid chromosome number may be a mistake. Böcher may have cultivated the plant and counted the haploid number (in PMC) as 24, as was his usually applied technique, and Gjærevoll may have (automatically) assumed it as the diploid count or used the common notation of '2n='. Ryvarden (pers. comm.) does not remember how it was done. Böcher vouchered his counts but the vouchers are stowed away and nearly impossible to find (pers. comm. from Copenhagen herbarium). A hexaploid number is consistent with both D. norvegica and D. arctogena. There are too many doubts as concern this 'entity' to give it any status at present. (Elven)


[Ser. Nivales Tolm. in Kom. (1939), Fl. SSSR 8: 415.]
The Draba nivalis aggregate (D. kamtschatica, D. lonchocarpa, D. nivalis, D. palanderiana)

Comments:

(1) The four species listed here seem to constitute a group of comparatively close low-ploid relatives. They are therefore entered as a D. nivalis aggregate. The aggregate also contains several non-arctic species, e.g., the Cordilleran D. porsildii G.A. Mulligan that reaches north to Yukon Terr. and EC Alaska. (Elven)
53.23.19  Draba nivalis Lilj. (1793), Utkast Sv. Fl. 236 (1792).

S

2n= 16 (2x).



2nD Heilborn (1927 Norw); Löve & Löve (1956b N Eur); Jørgensen et al. (1958 Grl); Zhukova (1965b NE As); Knaben (1966a Norw); Knaben & Engelskjøn (1967 Norw); Johnson & Packer (1968 Ala); Mulligan & Porsild (1969b Can); Zhukova & Petrovsky (1971, 1984 NE As); Zhukova & Tikhonova (1971, 1973 NE As); Mulligan (1974b N Am); Brochmann et al. (1993 Sb).

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:
53.23.20  Draba lonchocarpa Rydb. (1900), Mem. New York Bot. Gard. 1: 181.

S

2n= 16 (2x).



2nD Taylor & Mulligan (1968 N Am); Mulligan (1974b N Am); Zhukova & Petrovsky (1984 NE As).

G SIB RFE ALA?

Comments:

(1) Löve & Löve (1975) refer the American chromosome counts (and distribution) to subsp. kamtschatica, but this taxon is absent from the areas from where the counts are reported whereas D. lonchocarpa s. str. is present there. Draba lonchocarpa and D. kamtschatica are quite clearly different and should be recognised as different species. The question mark for Alaska is due to reports from Brooks Range (in ALA) that should be confirmed as coming from this species. The subspecies of Mulligan (1974) are insufficiently characterised and could perhaps be reduced to varieties, and only subsp./var. lonchocarpa seems to reach the Arctic. (Elven)


53.23.21  Draba kamtschatica (Ledeb.) N. Busch (1919), Fl. Siber. Orient. 3: 329.

B D. frigida Turcz. var. (b) kamtschatica Ledeb. (1842), Fl. Ross. 1: 150.

S D. lonchocarpa Rydb. subsp. kamtschatica Calder & Taylor (1965), Canad. J. Bot. 43: 1393.

2n= 16 (2x).

2nD Zhukova & Petrovsky (1984 NE As).

G SIB RFE

Comments:

(1) The type material (LE) differs appreciably from American material of D. lonchocarpa and we would go for status as separate species. (Petrovsky, Elven & Grunt)


53.23.22  Draba palanderiana Kjellm. in Nordenskiöld (1883), Vega Exp. Vetensk. Iaktt. 2: 45.

S D. caesia auct., non Adams (1817).

2n= (1) 16 (2x). (2) 32 (4x).

2nD (1) Mulligan in Löve & Löve (1975 N Am); see comments. (2) Zhukova (1968 Sib); Zhukova & Tikhonova (1971 Sib); Zhukova & Petrovsky (1984 NE As).

G RFE ALA CAN

Comments:

(1) The diploid chromosome number reported by Mulligan in Löve & Löve (1975) should not be accepted without confirmation by voucher control. There are several closely related diploid 'Nivales' species. (Elven)
[Ser. Cinereae Tolm. in Kom. (1939), Fl. SSSR 8: 419.]
The Draba cinerea aggregate (D. arctica, D. cinerea, D. oblongata, D. ovibovina, D. parvisiliquosa)

Comments:

(1) This is a 'natural' group of related taxa and is proposed entered as a D. cinerea aggregate. They are all polyploid (4x-10x) and it is probable (or at least possible) that many of the same original diploid genomes are present in many of the polyploids. The relationships between the taxa accepted here are, however, still uncertain. Draba cinerea, D. arctica and D. oblongata seem to represent three 'major' and fairly distinct species. Draba parvisiliquosa could perhaps be included in an extended D. cinerea. Draba ovibovina seems to be most closely related to D. arctica but is recorded with more or less constant differences from that species and at a different ploidy levels. All of them are therefore tentatively proposed accepted as species. (Elven)
53.23.23  Draba cinerea Adams (1817), Mém. Soc. Imp. Naturalistes Moscou 5: 103.

S

2n= (1) 32 (4x). (2) 48 (6x). (3) 56 (7x). (4) 64 (8x).



2nD (1) Petrovsky's draft checklist; see comments. (2) Heilborn (1927 **, 1941 Grl); Böcher (1966a Grl); Mulligan (1971 Can); Zhukova & Petrovsky (1984 NE As); Murray & Kelso (1997 Ala). (3) Petrovsky's draft checklist; see comments. (4) Mulligan (1971 Can); Zhukova & Petrovsky (1984? Sib); see comments.

G RUS SIB RFE ALA? CAN GRL

Comments: See also D. parvisiliquosa.

(1) The specimens available for typification (Moscow) do not fit very well the current concept of the species. A photograph of the type furnished by Dave confirms this; the specimen does not fit any arctic Draba I am familiar with. There may therefore be problematic to reach an unambiguous typification which could confirm the current usage of the name 'cinerea'. (Murray & Elven)

(2) In the N Atlantic areas, D. cinerea is comparatively southern. It occurs in S Greenland but not in the north. In the Nordic area it is restricted to the North Boreal parts of the mainland and is replaced by D. arctica in Svalbard. My experiences from Alaska and arctic Canada are similar; in the arctic islands I found D. arctica and D. oblongata but no D. cinerea as we define it; it occurred in boreal Alaska. The material from other arctic areas should be critically checked. (Elven)

(3) In view of the very varied concept of D. cinerea applied through time and across regions, all chromosome numbers except the well proved hexaploid should be checked thoroughly against vouchers. (Elven)


53.23.24  Draba parvisiliquosa Tolm. (1932), Tr. Bot. Mus. 24: 271.

S

2n= 48 (6x).



2nD Zhukova & Petrovsky (1984 NE As).

G SIB RFE GRL?

Comments:

(1) This species is insufficiently characterised at present. There is, however, a morphological variation in boreal and southern arctic D. cinerea s. lat. that may merit taxonomic recognition. Plants more or less fitting the description of D. parvisiliquosa also occur in non-arctic N Fennoscandia (D. 'cinerea' in other areas here) and in S Greenland. In Fennoscandia, they also differ in some molecular markers (Andersen in prep., Andersen et al. 1999). But, when plants that may be named as D. parvisiliquosa occur throughout a major part of the range of D. 'cinerea' (Greenland, Fennoscandia, Siberia, Russian Far East) we should consider whether they are different at level of species. (Elven)



WARNING! Might be merged with D. cinerea.
53.23.25  Draba arctica J. Vahl in Hornem. (1840), Fl. Dan. 13, 39: t. 2294.

S D. cinerea auct., non Adams (1817).

T Svalbard: Bellsound, [1838-1839], J.Vahl (C? S? O?), 'lectotype' indicated by Böcher (1966: 14), but not formally selected.

Comments:

(1) Morphologically and cytologically well separated from D. cinerea s. str. There are also consistent differences in genetic markers (Brochmann et al. 1992a, 1992b, 1993; Andersen et al. 1999; Andersen in prep.). In recent decades this species has been well understood in Greenland, Svalbard and Russia   Siberia, but in North America it was still included by implication in D. cinerea by Porsild & Cody (1980), Rollins (1993) and Cody (1996) and is not separated by Scott (in prep., Flora of the Canadian Arctic Archipelago). The North American distribution is therefore much less well known than in other areas.

I have tentatively included as subspecies the Greenland entities described and accepted by Ekman and Böcher. As for subsp. ostenfeldii, both Böcher (1966) and Andersen et al. (1999) found consistent differences from D. arctica s. str. that may justify treatment as a subspecies even if they are sympatric and both are decaploid. Species rank is out of the question for these 'close' entities and there is no ecological difference that could justify ecotypical varieties. (Elven)

(2) As to typification. Böcher (1966) cited one specimen in C and two in S. In addition there are relevant specimens in O. No formal lectotypification seems to have been made. The specimens in O all represent D. arctica as we consider it. In S, one of the sheets is from Svalbard: "Spitsbergen. misit auctor" (J. Vahl). This is clear D. arctica in our concept and would also be a natural choice as lectotype. The second sheet contains three specimens, two to the left from Svalbard ("Spitsbergen"   D. arctica), one to the right from Greenland (D. cinerea). (Elven)
53.23.25.1  Draba arctica J. Vahl subsp. arctica

S

2n= 80 (10x).



2nD Heilborn (1927 Sb, as D. cinerea); Flovik (1940 Sb, as D. cinerea); Jørgensen et al. (1958 Grl); Böcher (1966a Sb); Holmen in Böcher (1966a Grl); Mulligan (1971a N Am); Zhukova et al. (1973 Sib); Zhukova & Tikhonova (1973 Sib); Brochmann et al. (1993 Sb)

G NOR RUS? SIB RFE ALA CAN GRL

Comments:
53.23.25.2  Draba arctica J. Vahl subsp. ostenfeldii (E. Ekman) Böcher ex J.T. Kartesz & K.N. Gandhi (1992), Phytologia 72, 2: 84.

B D. ostenfeldii E. Ekman (1930), Sv. Bot. Tidskr. 23: 491, non D. x ostenfeldii O.E. Schulz (1927).

S

2n= 80 (10x).



2nD Böcher (1966c Grl).

G GRL


Comments:
53.23.26  Draba ovibovina (E. Ekman) E. Ekman (1941), Sv. Bot. Tidskr. 35: 135.

B D. ostenfeldii E. Ekman var. ovibovina E. Ekman (1929), Sv. Bot. Tidskr. 23: 493.

S

2n= 48 (6x).



2nD Jørgensen et al. (1958 Grl).

G GRL


Comments:

(1) This hexaploid entity was described as a variety of the otherwise decaploid D. ostenfeldii (D. arctica subsp. ostenfeldii). It might be a more minor race of D. cinerea. Further investigations are needed before it can be accepted. (Elven)



WARNING! Should perhaps be reduced to synonymy within D. cinerea.
53.23.27  Draba oblongata R. Br. [(1819), nom. nud., List Pl. Ross. Voy. App. 143] ex DC. (1821), Syst. Nat. 2: 342.

S D. groenlandica E. Ekman (1929), Sv. Bot. Tidskr. 23: 4 [or 486]; D. arctica J. Vahl subsp. groenlandica (E. Ekman) Böcher in Böcher et al. (1968), Grønlands flora 92.

T *** (K), see Mulligan (1974a).

2n= 64 (8x).

2nD Heilborn (1941 Grl, count probably made on type material of D. groenlandica); Holmen (1952 Grl); Jørgensen et al. (1958 Grl); Böcher (1966a Grl); Zhukova (1967a Sib); Mulligan (1971a, 1974a N Am); Zhukova & Petrovsky (1984 NE As).

G NOR? RUS SIB RFE ALA? CAN GRL

Comments:

(1) The name D. oblongata was for a long time misapplied for a yellow-flowered species (D. micropetala Hook.) but is now, after the elucidation by Mulligan (1974a), well established as the valid name for what previously was known as D. groenlandica. The affinity of the species is not entirely uncertain. It combines features from ser. Cinereae and ser. Rupestres and is an argument for lumping these series.

The question mark for Norway relates to recently discovered collections from northern Spitsbergen that resemple D. oblongata in most characters. The site is inaccessible and it is difficult to base a firm conclusion on the specimens available. (Elven)
[Ser. Rupestres Tolm. in Kom. (1939), Fl. SSSR 8: 421.]
The Draba norvegica aggregate (D. arctogena, D. norvegica)

Comments:

(1) Two of the three listed species in ser. Rupestres (D. norvegica and D. arctogena) are very closely related and constitute a natural aggregate. The third   D. baicalensis   is of much more uncertain affinity. (Elven)
53.23.28  Draba norvegica Gunnerus (1772), Fl. Norveg. 2: 106.

S D. hirta L. (1759) nom. rejic. prop. p.p.; D. rupestris R. Br. in W.T. Aiton (1812), Hortus Kew., ed. 2, 4: 91 (1812).

T *** (TRH).

2n= 48 (6x).

2nD Heilborn (1927 Norw); Flovik (1940 Sb); Heilborn (1941 Norw, Grl); Löve & Löve (1956b N Eur); Böcher (1966a Grl); Knaben (1966a Norw); Knaben & Engelskjøn (1967 Norw); Mulligan & Cody (1968 N Am); Mulligan (1970 N Am); Engelskjøn (1979 Sb, Bear Island); Brochmann et al. (1993 Sb).

G ICE NOR RUS RFE? CAN GRL

Comments:

(1) Closely related to D. arctogena and intermediates are reported in Greenland (Böcher 1966a). They may represent N Atlantic ('norvegica') and Greenland   N America   NE Siberia ('arctogena') parts of one polymorphic species, i.e., subspecies.

The records from Russian Far East must be thoroughly checked as there otherwise is no reliable record of this species from NW Canada, Alaska or Siberia. (Elven)
53.23.29  Draba arctogena (E. Ekman) E. Ekman in Grøntved (1936), Vasc. Pl. Arct. N. Amer. 83.

B D. groenlandica E. Ekman var. arctogena E. Ekman (1929), Sv. Bot. Tidskr. 23: 489.

S

T *** (S).



2n= 48 (6x).

2nD Heilborn (1941 Grl); Holmen (1952 Grl); Zhukova & Petrovsky (1971, 1984 NE As); Zhukova et al. (1973 NE As).

G NOR? RFE ALA CAN GRL

Comments:

(1) The question mark for Norway is based on specimens from NW Spitsbergen (Svalbard, TRH) that fit much better with Greenland D. arctogena than with the comparatively frequent D. norvegica in Svalbard. (Andersen & Elven)
53.23.30  Draba baicalensis Tolm. in Kom. (1939), Fl. SSSR 8: 421, 650.

S ?D. yukonensis A.E. Porsild (1975), Natl. Mus. Canada Publ. Bot. 4: 37.

2n=

2nD


G RFE

Comments:

(1) Validity of the species to be verified. Might be the same as or very close to the American D. yukonensis (acc. to A. Berkutenko) which here is given as a possible synonym. Draba yukonensis does not reach the Arctic. (Elven)
[Ser. Hirtae Tolm. in Kom. (1939), Fl. SSSR 8: 427.]
The Draba glabella aggregate (D. glabella, D. juvenilis, D. praealta)

Comments:

(1) Three of the five listed species seem to constitute an aggregate (D. glabella, D. juvenilis and D. praealta). The two last ones are much more uncertain. (Elven)
53.23.31  Draba glabella Pursh (1814), Fl. Amer. Sept. 2: 434.

S D. hirta L. (1759), Syst. Nat. 2: 350, nom. rejic. prop. p.p.; D. daurica DC. (1821), Syst. Nat. 2: 350.

2n= (1) 64 (8x). (2) c. 75. (3) 80 (10x).

2nD (1) Heilborn (1927 Grl, 1941 Grl & Norw); Böcher & Larsen (1950 Grl); Löve & Löve (1956b N Eur, but see comments); Holmen in Jørgensen et al. (1958 Grl); Böcher (1966a Can & Grl); Holmen in Böcher (1966a Grl); Knaben & Engelskjøn (1967 Norw); Zhukova (1967a Sib); Mulligan (1970 N Am); Zhukova & Tikhonova (1971, 1973 Sib); Zhukova & Petrovsky (1984 Sib); Brochmann et al. (1993 Sb); Murray & Kelso (1997 Ala). (2) Rollins (1966 Ala); see comments. (3) Heilborn (1927 Grl); Jørgensen et al. (1958 Grl); Johnson & Packer (1968 Ala).

G ICE? NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) As to typification of D. hirta Linnaeus. The material available for typification is LINN 823.12 with two specimens. The left-hand plant is marked "lapp" (Lapland, i.e., N Scandinavia), is in acceptable condition and may belong to D. glabella as currently understood. The right-hand plant is in very bad condition and probably belongs to D. norvegica as currently understood. However, the name has been applied and misapplied in too many meaning to have any current use. It is therefore, or at least should be, proposed rejected. (Elven)

(2) The deviant chromosome number of 2n=c.75 from Alaska (Rollins 1966) should be controlled by voucher. There seems, however, to be good evidence for the occurrence of two ploidy levels (2n=64 and 80) in this very widespread and polymorphic species. (Elven)

(3) The chromosome reports from Iceland (Löve & Löve 1956b, 1975) are enigmatic as the presence of this species in Iceland is discounted (Jalas & Suominen 1996, Kristinsson). The problem is that no octoploid Draba is currently known from Iceland. Vouchers of Löve counts are notoriously difficult to find. (Elven)
53.23.32  Draba juvenilis (Pohle) Kom. (1914), Feddes Repert. 13: 163.

B Draba hirta L. var. juvenilis Pohle (1925), Drabae Asiat. 36.

S D. longipes Raup (1934), Contr. Arnold Arbor. 6: 165.

2n= (1) 48 (6x). (2) 64 (8x).

2nD (1) Zhukova & Petrovsky (1972, 1984 NE As, as D. juvenilis). (2) Mulligan (1970 N Am, as D. longipes).

G SIB RFE ALA CAN

Comments:
53.23.33  Draba praealta E.L. Greene (1898), Pittonia 3: 306.

S

2n= 56 (8x with x=7?).



2nD Mulligan (1966 N Am).

G RFE ALA

Comments:
53.23.34  Draba prozorovskii Tolm. (1930), Compt. Rend. Acad. Sci. URSS, Ser. A, 7: 173.

S

2n=



2nD

G SIB


Comments:

(1) Validity of species must be verified. It is reported as very scattered in N and NE Siberia and was also once described by Trautvetter (in sched.) as a variety of D. incana. (Elven)



WARNING! In need of much more documentation before we can accept it.
53.23.35  Draba chamissonis G. Don (1831), Gen. Syst. 1: 184.

S

2n=



2nD

G RFE ALA

Comments:

(1) Validity and affinity of species must be verified. The species is tentatively included by Petrovsky in this series; by Tolmachev (1939, 1975) it was placed in ser. Nivales. Tolmachev (1975) mapped it from five widely scattered sites in NE Asia and Hultén (1968) has two sites in NW Alaska, in addition to two sites in easternmost Chukchi Peninsula. The Chukchi sites are not mentioned or mapped by Tolmachev. This is really strange as Hultén (1968) stated that the species was described from Chukchi Peninsula. It is impossible to get any consistent impression of this species from the descriptions and few comments I have available. (Elven)



WARNING! Very little is known about this entity; in danger of exclusion.
[Ser. Incanae Tolm. in Kom. (1939), Fl. SSSR 8: 432.]
The Draba incana aggregate (D. cana, D. incana)

Comments:

(1) The two listed species are closely related and constitute a 'natural' aggregate. (Elven)
53.23.36  Draba incana L. (1753), Sp. Pl. 643.

S

2n= 32 (4x).



2nD Heilborn (1927 Norw); Böcher & Larsen (1950 Grl); Löve & Löve (1956b N Eur); Jørgensen et al. (1958 Grl); Böcher (1966 Grl); Mulligan (1970 N Am); Laane (1971 Norw).

G ICE NOR RUS CAN GRL

Comments:

(1) The species, even in its original (European) meaning, is heterogeneous. There are arguments for separation of a W and C European subspecies (D. incana L. s.str.) and a much more widespread northern subspecies. The affinities of the latter to D. lanceolata Royle and D. cana should be investigated. (Elven)


53.23.37  Draba cana Rydb. (1902), Bull. Torrey Bot. Club 29: 241.

S D. valida Goodding (1904), Bot. Gaz. 37: 35; D. breweri S. Wats. (1888, Proc. Amer. Acad. Arts 23: 260) var. cana (Rydb.) Rollins (1993), Harvard Papers Bot. 44: 46; D. lanceolata auct., non Royle (1839).

2n= 32 (4x).

2nD Böcher (1966a Grl, as D. lanceolata); Mulligan (1966, 1971a N Am).

G ALA CAN GRL

Comments:

(1) Draba lanceolata Royle (1839, Illustr. Bot. Himal. Mts. 1: 72) was included in Petrovsky's draft but is here excluded. In the Arctic, D. lanceolata is only reported from North America and Greenland, but all these reports are now referred to D. cana Rydb. or alternatively D. breweri var. cana (Rollins 1993). Draba lanceolata Royle is probably an Asiatic species which do not reach the Arctic proper. The reported chromosome numbers, 2n=32 (4x, Podlech & Dieterle 1969 from Afghanistan, Krogulevich 1971 from the Baical area) and 48 (6x, from where?) are probably all non-arctic.

Due to confusion with D. lanceolata Royle, only Greenland   N American chromosome counts are accepted. Rollins (1993) reduced D. cana to a variety of the otherwise American D. breweri S. Wats. This has not been followed here. (Elven)


[Ser. Boreales Pohle (19**), ***.]
53.23.38  Draba borealis DC. (1821), Syst. Nat. 2: 342.

S Draba luteola auct., non E.L. Greene (1899).

2n= (1) 64 (8x). (2) 80 (10x).

2nD (1) Zhukova & Petrovsky (1984 NE As). (2) Löve & Löve (1975); see comments.

G SIB RFE ALA CAN

Comments:

(1) Löve & Löve (1975) reported only the decaploid number (2n=80) for this species. They reported it as Beringian only, but a majority of the counts they listed are from Greenland. They seem to have confused D. borealis with D. arctica (or D. aurea). The decaploid numbers reported should therefore be checked throrougly against available vouchers before acceptance. (Elven)
[Ser. Repentes Tolm. in Kom. (1939), Fl. SSSR 8: 443.]
53.23.39  Draba sibirica (Pall.) Thell. (1906), Gatt. Lepidium, 318 in nota 2.

S Lepidium sibiricum Pall. (1776), Reise 3: 34.

2n= 16 (2x).

2nD Böcher (1966 garden origin, 1974 Grl); Mulligan in Porsild (1967, 1969b N Am?).

G RUS SIB GRL

Comments:

(1) Löve & Löve (1975) report the arctic plants of D. sibirica as a subsp. arctica Böcher ***). (Elven)

(2) A related but distinct species   D. ogilviensis Hultén (1966), Bot. Not. 119: 315   is described as an endemic from Ogilvie Mts on the Alaskan/Canadian border, Richardson Mts and Wrangell - St. Elias Mts, south of our arctic boundary. See Murray & Parker (1999, Canad. Field-Natural. 113: 659-662). (Elven)


[Ser. Nemorosae Tolm. in Kom. (1939), Fl. SSSR 8: 449.]
53.23.40  Draba nemorosa L. (1753), Sp. Pl. 643.

S

2n= 16 (2x).



2nD Packer (1964 N Am); Mulligan (1966 N Am); Zhukova et al. (1973 Sib).

G RUS SIB RFE ALA CAN

Comments:
[Ser. Aureae V.V. Petrovsky (19**), ***.]
53.23.41  Draba aurea Vahl ex Hornem. (1806), Fors. Oecon. Plantel., ed. 2, 599.

S Draba luteola E.L. Greene (1899), Pittonia 4: 19.

2n= (1) 40+1. (2) 64. (3) 72. (4) 74. (5) 76. (6) 82.

2nD (1) Rollins (1993). (2) Rollins (1993). (3) Böcher (1966 Grl, interpreted as possible 72+4). (4) Mulligan (1966 N Am). (5) See (3). (6) Rollins (1993).

G CAN GRL

Comments:

(1) The many and very deviating chromosome counts should be checked against sources and vouchers. (Elven)
Ser. ???
The Draba incerta aggregate (D. exalata, D. incerta)

Comments:

(1) Löve & Löve (1975) synonymized D. incerta Payson with D. exalata E. Ekman, which in turn is synonymized by some other authors (e.g., Hultén 1968) with D. ruaxes Payson & St. John and D. ventosa A. Gray (1874, Amer. Natural. 8: 212). This species or species group (aggregate) combines features from ser. Pilosae and ser. Alpinae. Even if it was revised by Mulligan (1971b), it is probably still in need of some study. (Elven)
53.23.43  Draba incerta Payson (1917), Amer. J. Bot. 4: 261.

S

2n= 112 (14x).



2nD Mulligan (1966, 1972 N Am).

G ALA CAN

Comments:
53.23.44  Draba exalata E. Ekman (1932), Sv. Bot. Tidskr. 25: 489.

S

2n= (1) 72 (9x). (2) 112 (14x).



2nD (1) ? (2) ?

G ALA


Comments:

(1) Possibly synonymous with D. incerta and to be excluded. The species is not mentioned by Rollins (1993). However, Cody (1996) reports D. incerta only north to SE Alaska and to British Mts. in Yukon Territory, which must mean that he excludes from D. incerta the plants reported as D. exalata by Hultén (1968) from Seward Peninsula. (Elven)


Ser. 'Drabella' ???
53.23.45  Draba crassifolia Graham (1829), Edinburgh New Philos. J. 7: 182.

S

2n= 40 (4x or 8x with x=10 or 5).



2nD Heilborn (1941 Grl); Mulligan (1966, 1975 N Am); Engelskjøn & Knaben (1971 N Norw).

G RFE CAN GRL

Comments:

(1) The probably related D. albertina E.L. Greene (1901), Pittonia 4: 312, is Cordilleran and does not reach the Arctic. It is reported as di  and tetraploid (2n=12, 24) with base number x=6. (Elven)


53.23.46  Draba stenoloba Ledeb. (1842), Fl. Ross. 1: 154.

S

2n= 40 (4x or 8x with x=10 or 5).



2nD Mulligan (1970 N Am).

G ALA CAN



Comments:


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