Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 73, 98




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Sect. Orobia Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 73, 98.

S Sect. Diphragmae Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: ***

Comments:

(1) Comparatively many of the entities given below are weakly defined and should probably be merged into more widely defined species. For very many of them we also have little information. Russians should look into 'their' species and North Americans into 'theirs'. I have proposed comparatively few changes yet. (Elven)


The Oxytropis adamsiana aggregate (O. adamsiana, O. uschakovii, O. vassilczenkoi)

Comments:

(1) The aggregate disappears if all three are merged in an extended O. adamsiana. (Elven)
60.6.15  Oxytropis adamsiana (Trautv.) Jurtz. (1959), Bot. Mater. Gerb. Bot. Inst. AN SSSR 19: 248.

B O. strobilacea Bunge var. adamsiana Trautv. (1877), Tr. St. Petersb. Bot. Sada 5, 1[Pl. Sib. Bor.]: 40.

S

Comments:



(1) The two proposed subspecies are parapatric, subsp. adamsiana from Taimyr to Kharaulakh, subsp. janensis from Kharaulakh eastwards. Their status should be evaluated. (Elven)
60.6.15.1  Oxytropis adamsiana (Trautv.) Jurtz. subsp. adamsiana

S

2n= 32 (4x).



2nD Yurtsev & Zhukova (1968); Zhukova & Petrovsky (1971).

G SIB


Comments:
60.6.15.2  Oxytropis adamsiana (Trautv.) Jurtz. subsp. janensis Jurtz. (1959), Bot. Mater. Gerb. Bot. Inst. AN SSSR 19: 261.

S

2n= 32 (4x).



2nD ?

G SIB


Comments:
60.6.16  Oxytropis uschakovii Jurtz. (1986), Arkt. Fl. SSSR 9,2: 104, 178.

S

T Russian Far East: Insula Wrangelii, sinus Somnitelnaja (Dubius), regio ripae dextrae fl. Somnitelnaja adjacens prope montes, 02.08.1065, leg. B.A.Jurtzev (LE) holotype.



2n= 32 (4x).

2nD ?


G RFE

Comments:

(1) A local endemic of Wrangel Island, previously considered as part of O. adamsiana. Status as separate species? (Elven)

WARNING! Might be reduced to a race of O. adamsiana.
60.6.17  Oxytropis vassilczenkoi Jurtz. (1959), Bot. Mater. Gerb. Bot. Inst. AN SSSR 19: 262.

S

Comments:



(1) Previously and also more recently considered as part (race) of O. adamsiana, see Kozhevnikov (1980, Bot. Zhurn. 65, 3). It replaces O. adamsiana east of Kolyma as would be natural for a geographical race. The two subspecies accepted by Yurtsev are nearly fully sympatric and should be very critically viewed. (Elven)

WARNING! Might be reduced to a race of O. adamsiana; subspecies might be merged.
60.6.17.1  Oxytropis vassilczenkoi Jurtz. subsp. vassilczenkoi

S

2n= (1) 32 (4x). (2) 48 (6x).



2nD (1) Sokolovskaya (1968); Zhukova (1969); Zhukova & Tikhonova (1971).

G RFE


Comments:
60.6.17.2  Oxytropis vassilczenkoi Jurtz. subsp. substepposa Jurtz. (1986), Fl. Arct. URSS 9, 2: 102, 178.

S

T Russian Far East: Terra Tschuktschorum occidentalis, in litore orientali sinus Czaunensi prope pagum Valkumei, 06.07.1980, leg., T.G. Polozova & B.A. Yurtsev (LE) holotype.



2n= (1) 32 (4x). (2) 48 (6x).

2nD ?


G RFE

Comments:


60.6.18  Oxytropis arctica R. Br. (1823), Chloris Melvill. 20.

S

T Canada: Melville Island, leg. Mr. Beverley (K) lectotype, selected by Welsh (1995. Great Basin Natural. 55: ***).



Comments:

(1) What about Chloris Melvilliana (1823)? Might have priority. (Elven)

(2) The Asian and North American parts are probably different enough to merit rank as two major races, i.e. subspecies. (Elven)
60.6.18.1  Oxytropis arctica R. Br. subsp. arctica

S

2n= (1) c. 80 (10x). (2) 96 (12x).



2nD (1) Yurtsev's draft. (2) Mosquin & Hayley (1966, 2n=c.96); Löve & Löve in Löve (1975d).

G ALA CAN

Comments:
60.6.18.2  Oxytropis arctica R. Br. subsp. taimyrensis Jurtz. (1959), Bot. Mater. Gerb. Bot. Inst. AN SSSR 19: 239.

S O. karga Sapozhn. ex Polozh. (1956), Sist. Zam. Gerb. Tomsk. Univ. 79-80: 2; O. taimyrensis (Jurtz.) Á. Löve & D. Löve (1976), Bot. Not. 128: 516.

2n= 64 (8x).

2nD Ledingham & Reever (1963).

G SIB

Comments:


60.6.19  Oxytropis koyukukensis A.E. Porsild (1951), Canad. Field-Natural. 65: 78.

S O. arctica R. Br. var. koyukukensis (A.E. Porsild) S.L. Welsh (***), 'Legumes of Alaska 2': 280.

2n= 48 (6x).

2nD Holmen (1962).

G ALA

Comments:



(1) The differences given by Hultén (1968, Ark. Bot.) between O. koyukukensis and O. arctica are probably too few and too small to justify rank as species. (Elven)

WARNING! Should probably be merged with O. arctica, perhaps as a race.
60.6.20  Oxytropis tichomirovii Jurtz. (1986), Fl. Arct. URSS 9, 2: 128, 180.

S

T Siberia: Taimyr, districtus Avamensis, regio ripae dextrae fl. Volczja adjacens, 10.07.1964, leg. G.Lebedeva (LE) holotype.



2n= 16 (2x).

2nD ?


G SIB

Comments:

(1) A plant with a combination of characters of O. arctica and O. nigrescens, probably a hybridogenous species. (Yurtsev)

(2) This entity occurs entirely within the range of both O. arctica and O. nigrescens. The ploidy level is indicated as diploid   as in O. nigrescens   and markedly different from the octoploid level indicated for O. arctica (subsp. taimyrensis). A diploid hybrid between a diploid and an octoploid seems strange. Perhaps O. tichomirovii rather should be merged with O. nigrescens or at least moved to that affinity? (Elven)



WARNING! Might be merged with O. nigrescens, see comment below.
60.6.21  Oxytropis semiglobosa Jurtz. (1964), Nov. Sist. Vyssh. Rast. 1: 217.

S O. ajanensis (Regel & Tiling) Bunge subsp. semiglobosa (Jurtz.) N.S. Pavlova (1989), Sosud. Rast. Sovet. Dal'nego Vostoka 4: 269.

2n= 16 (2x).

2nD Yurtsev & Zhukova (1972).

G RFE

Comments: More information needed.


60.6.22  Oxytropis ochotensis Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 95.

S

2n= (1) 48 (6x). (2) 64 (8x).



2nD ?

G SIB RFE

Comments: More information needed.
60.6.23  Oxytropis maydelliana Trautv. (1879), Acta Horti Petrop. [=Glavn. Bot. Sada] 6: 16.

S

T Russian Far East: in tractu fluvium Anadyr inter et latus meridionale montium jugi a fluvio hoc septentrionem versus siti, 1869, leg. Baron G. von Maydell (LE) holotype?



Comments:

(1) Gillett et al. (1999 onwards) merge subsp. maydelliana and subsp. melanocarpa without further comment. The main diagnostic character (inflorescence indumentum) seems to hold and I propose that we keep two races, one amphi-Beringian east to Mackenzie River and one North American from Mackenzie eastwards. Intermediates occur in the Mackenzie River area but that is to be expected among subspecies. (Elven)


60.6.23.1  Oxytropis maydelliana Trautv. subsp. maydelliana

S

2n= 96 (12x).



2nD Löve & Löve (1975) list six counts, five as arctic, all for subsp. maydelliana.

G RFE ALA CAN

Comments:
60.6.23.2  Oxytropis maydelliana Trautv. subsp. melanocephala (Hook.) A.E. Porsild (1966), Bull. Natl. Mus. Can. 216: 47.

S O. campestris (L.) DC. var. melanocephala Hook. (1831), Fl. Bor.-Amer. 1: 147.

T [Described from Canada: "Bear Lake to the Arctic Shores and Islands".]

2n=


2nD

G CAN


Comments:
The Oxytropis campestris aggregate (O. beringensis, O. hyperborea, O. johannensis, O. jordalii, O. katangensis, O. leucantha, ?O. roaldii, O. sordida, ?O. terrae-novae, O. varians)

Comments:

(1) The treatments of this aggregate diverge widely. Yurtsev considers it as a group of (more or less) well-defined species, most of them allopatric or parapatric. Gillett et al. (1999 onwards) reduce the entities that reach the Arctic Archipelago to varieties of a wide O. campestris. European views have mainly been the same but with usage of subspecies instead of variety. My (restricted) experiences in N Europe and Alaska-Canada more point towards subspecies than separate species for most of thise entities I have seen. (Elven)
60.6.24  Oxytropis leucantha (Pall.) Bunge (1874), Mém. Acad. Imp. Sci. St.-Pétersb., ser. 7, 22, 2: 111 [ememd. Welsh (1972), Taxon 21: 155].

B Astragalus leucanthus Pall. (1800), Spec. Astragal. 59.

S

Comments:



(1) Hultén (1947) considers O. leucantha as related to the O. borealis DC. group. However, many early sources relate it to O. campestris s. lat. See, however, Hultén (1968, Fl. Alaska p. 667). (Elven)

(2) The two subspecies given in Yurtsev's draft are parapatric. We need more information on how well they are separated. (Elven)


60.6.24.1  Oxytropis leucantha (Pall.) Bunge subsp. subarctica Jurtz. (1986), Fl. Arct. URSS 9, 2: 119, 180.

S

T Siberia: Jacutia, ad Lenam inferiorem, fl Czubukulach, 29.07.1957, leg. V.V. Petrovsky & B.A. Yurtsev (LE) holotype.



2n= (1) 48 (6x). (2) 96 (12x).

2nD ?


G SIB

Comments:


60.6.24.2  Oxytropis leucantha (Pall.) Bunge subsp. tschukotcensis Jurtz. (1986), Fl. Arct. URSS 9, 2: 120, 180.

S O. dorogostajskyi auct., non Kuzen. (1920).

T Russian Far East: Terra Tschuktschorum occidentalis, pars australi-occidentalis planitiei montium Czukotensis, regio montana ripae dextrae fl. Palavaam adjacens, in 6 km a pago Palavaam adverso flumine, 08.07.1973, leg. S.A. Balandin, V.Yu. Raszhivin, B.A. Jurtzev et al. (LE) holotype.

2n= (1) 48 (6x). (2) 64 (8x). (3) 96 (12x).

2nD ?

G SIB RFE



Comments:
60.6.25  Oxytropis jordalii A.E. Porsild (1951), Canad. Field-Natural. 65: 77.

S O. campestris (L.) DC. subsp. jordalii (A.E. Pors.) Hultén (1968), Ark. Bot., ser. 2, 7, 1: 79; O. leucantha (Pall.) Bunge subsp. jordalii (A.E. Porsild) Jurtz. (1980), Byull. Mosk. Obshch. Ispyt. Prir., Biol. 85, 6: 104.

2n= (1) 16 (2x). (2) 96 (12x).

2nD (1) Yurtsev's draft, misprint? (2) Zhukova & Petrovsky (1972).

G RFE? ALA CAN

Comments:

(1) Considered by North American botanists (e.g., Cody 1996) an Alaskan and NW Canadian mountain race of O. campestris. We should adopt that view, i.e., as subsp. jordalii. Hultén (1968, Ark. Bot. & Fl. Alaska) considers it as closely related to and an American counterpart to the 'sordida' entity. He also includes a site in Chukchi Peninsula. (Elven)

WARNING! Will most probably be reduced to a subspecies of O. campestris.
60.6.26  Oxytropis varians (Rydb.) K. Schum. (1903), Justs Bot. Jahresb. 19: 543.

B Aragallus varians Rydb. (1901), Bull. New York Bot. Gard. 2: 176.

S O. campestris (L.) DC. var. varians (Rydb.) Barneby (1952), Proc. Calif. Acad. Sci. 27: 253; O. hyperborea A.E. Porsild (1943), Sargentia 4: 53; O. campestris (L.) DC. subsp. gracilis (A. Nels.) Hultén (1968), Ark. Bot., ser. 2, 7, 1: 79 [basionym: Aragallus gracilis A. Nels. (1899), Erythea 7: 60].

T Yukon District: Lewes R., 28.06.1899, leg. Tarleton 33b (US) holotype.

2n= >60 (8x?).

2nD Johnson & Packer (1968 Ala, for O. gracilis).

G ALA? CAN

Comments:

(1) The combination O. varians (Rydb.) Hultén, made anew by Hultén (1947), Lunds Univ. Årsskr., n. f., avd. 2, 43, 1: 1107, is superfluous. (Elven)

(2) Oxytropis hyperborea is now synonymised with O. varians (or rather O. campestris subsp./var. varians by most current North American authors (e.g., Cody 1996, Gillett et al. 1999 onwards). I have followed this. Hultén (1947 p. 1108) summarised his opinion like this: "I can see no difference whatever between this plant [O. varians] and O. hyperborea as described by Porsild ... I must therefore regard Oxytropis hyperborea Pors. as a synonym of Aragallus varians Rydb." Oxytropis hyperborea is typified from Canada: NWT, Mackenzie R. Delta, E branch, 6855'N, 21.07.1934, leg. A.E. Porsild 7033 (CAN) holotype. Reported in Yurtsev's draft from ALA and CAN based on Porsild & Cody (1980). (Elven)

(3) Oxytropis gracilis / O. campestris subsp. gracilis (A. Nels.) Hultén (1968, Ark. Bot., ser. 2, 7, 1: 79) is synonymised with the 'varians' entity. The synonyms have been included as this is an accepted name in Hultén (1968). If the 'varians' entity is considered as a subspecies, as it probably should be, the name subsp. gracilis probably has priority. (Elven)

(4) About O. hyperborea: The record of 2n=c.60 (64) from Cape Thomson vicinity (Johnson & Packer 1968) could belong to this taxon, which could be considered to be a race of O. leucantha s. l. (Yurtsev)



WARNING! Will most probably be reduced to a subspecies of O. campestris.
60.6.27  Oxytropis katangensis Basil. (1924), Bot. Mater. Gerb. Bot. Inst. AN SSSR 5, 5: 3.

S O. schischkinii V.Vassil. (1948), Fl. SSSR 13: 66.

2n= 96 (12x).

2nD Ledingham & Reever (1963).

G SIB

Comments:



(1) A local species in the Jenisei area, known from only a few sites acc. to Fl. Arct. URSS 9, 2, map. 27 (p. 127). It keys out close to O. sordida and O. leucantha and should probably be included in synonymy with one of the subspecies of O. campestris. (Elven)

WARNING! Should probably be synonymised with a subspecies of O. campestris.
60.6.28  Oxytropis sordida (Willd.) Pers. (1807), Syn. Pl. 2: 332.

B Astragalus sordidus Willd. (1800), Sp. Pl. 3, 2: 1313.

S O. campestris (L.) DC. [(1802), Astragalogia 74; basionym: Astragalus campestris L. (1753), Sp. Pl. 761] subsp. sordida (Willd.) Hartm. fil. in Hartm. (1879), Handb. Skand. Fl., ed. 11, 305.

T [Described from Scotland and Norway.]

Comments:

(1) This is the major arctic Eurasiatic entity of the O. campestris aggregate. According to Fl. Arct. URSS, it is distributed from NE Norway (and Finland) eastwards to Russian Far East and perhaps across the Bering Strait (the 'barnebyana' entity). The map in Fl. Arct. URSS 9, 2 (map 24, p. 112) throws strong doubt on the applied subspecific (and perhaps specific) taxonomy. It is mapped as subsp. sordida more or less frequently eastwards to Jenisei and as scattered in interior Taimyr. Then a subsp. arctolenensis is mapped from three sites in lower Lena as its entire area. A third entity, subsp. schamurinii, is mapped from a single site in the Mys Svjatoy Nos area in Yana-Kolyma, from two sites in W Chukotka, and from Wrangel Island. The indicated close relative O. beringensis is mapped from one site in E Chukotka.

I would prefer to interpret the scattered E Siberian and Far East occurrences as slightly differentiated isolated populations and population groups of the 'sordida' entity rather than as taxa with their own rank. Similar isolated population groups are known many places elsewhere within the range of O. campestris s. lat. Recognition of every marginal population as a taxon of its own leads to confusion rather than elucidation. The deviating morphology should rather be commented on. (Elven)

WARNING! The 'species' should probably be reduced to a major race, i.e., subspecies, of O. campestris. The subspecies should probably be omitted.
60.6.28.1  Oxytropis sordida (Willd.) Pers. subsp. sordida

S

2n= 48 (6x).



2nD Jalas (1950 Finl); Sorsa (1963c Finl); Laane (1966 N Norw); Sokolovskaya (1970).

G NOR RUS SIB

Comments:
60.6.28.2  Oxytropis sordida (Willd.) Pers. subsp. arctolenensis Jurtz. (1986), Fl. Arct. URSS 9, 2: 114, 179.

S

T Siberia: Jacutia [arctica], ad Lenam inferiorem, prope stationem polarem "Sokol", 16.08.1956, leg. T. Polozova & B.A. Yurtsev (LE) holotype.



2n= 96 (12x).

2nD ?


G SIB

Comments: See comment above.


60.6.28.3  Oxytropis sordida (Willd.) Pers. subsp. schamurinii Jurtz. (1986), Fl. Arct. URSS 9, 2: 115, 179.

S

T Russian Far East: Insula Wrangelii, ad litus sini Somnitelnaja (Dubius), pedes declivii australis montis ad ripam dextram fl. Somnitelnaja, 31.07.1966, leg. V.F. Schamurin (LE) holotype.



2n= 96 (12x).

2nD ?


G SIB RFE

Comments: See comment above.


60.6.28.4  Oxytropis sordida (Willd.) Pers. subsp. barnebyana (Welsh) Jurtz. (1986), Fl. Arct. URSS 9, 2: 109.

B O. arctica R. Br. var. barnebyana Welsh (1968), Great Basin Natural. 28: 152.

S

2n=


2nD

G ALA


Comments: See also comment above.

(1) The 'barnebyana' entity that connects O. 'sordida' to North America and the report of a possible Chukotkan occurrence of the otherwise North American O. 'jordalii' should be compared. They may both only represent some intermediates or connection between the Asian and the North American side of the aggregate. See also the next 'species'. (Elven)


60.6.29  Oxytropis beringensis Jurtz. (1986), Fl. Arct. URSS 9, 2: 116, 179.

S

T Russian Far East: Peninsula Czukotskyi (Czukotensis), regio ripae sinistrae fl. Czegitun adjacens, in valle fl. Putukunei-Veem 1 km ab ostio, 21.07.1972, leg. V. Raszhivin, N. Sekretareva & B. Yurtsev (LE) holotype.



2n= 96 (12x).

2nD ?


G RFE

Comments:

(1) The consideration of this taxon known from two localities as a subspecies of O. sordida should be discussed; see Yurtsev et al. (1994). (Yurtsev)

(2) I can't suppress the suspicion that this 'entity' also is part of the transition between Asian ('sordida') and North American ('jordalii', 'varians/gracilis') major races of O. campestris. We should not accept it, neither as a species nor as a separate subspecies, without much more convincing justification. (Elven)


60.6.30  Oxytropis roaldii Ostenf. (1910), Chr. Vidensk. Selsk. Skr. 1909, 8: 54.

S O. campestris (L.) DC. var. roaldii (Ostenf.) S.L. Welsh (1991, Great Basin Natural. 51: ***; O. campestris (L.) DC. subsp. roaldii (Ostenf.) Cody ***; O. arctica auct., non R. Br. (1823).

T Yukon District: Herschel I., 13.07.1906, leg. Lindström (NY) holotype.

2n=


2nD

G ALA? CAN

Comments:

(1) Hultén (1947 p. 1106) related O. roaldii to O. arctica and did so also later (Hultén 1968, Fl. Alaska; 1968, Ark. Bot. where he included it together with O. coronaminis in O. arctica). Hultén seems to have had only 3-4 collections available whereas it now is recognised from numerous sites in N Alaska, N Yukon and NWT. It was recognised by Welsh (1991) as a variety of O. campestris and by Cody (19**) as a subspecies of the same. These opinions should be taken into account and evaluated. I will follow Cody and reduce it to an Alaskan   Canadian subspecies of O. campestris (and have moved it here in the sequence) if good arguments for its status as a species do not appear. (Elven)

(2) If we utilise the data of Hultén (1947), this entity also occurs in arctic Alaska (Teller in Sweard Peninsula). This si also indicated by Cody (1996). (Elven)

WARNING! Will probably be reduced to a subspecies of O. campestris.
60.6.31  Oxytropis terrae-novae Fern. (1928), Rhodora 30: 147.

S O. campestris (L.) DC. var. terrae-novae (Fern.) Barneby (1952), Proc. Calif. Acad. Sci. 27: 266; O. campestris (L.) DC. var. minor (Hook.) S.L. Welsh (1995), Great Basin Natural. 55: 277 [based on: O. uralensis taxon minor Hook. (1831), Fl. Bor.-Amer. 1: 146; type: Canada: Labrador, "O. uralensis gamma", leg. Morrison (K Hooker herb.) holotype, selected by Welsh (1995)]; O. johannensis (Fern.) Fern. (1928), Rhodora 30: 145 [basionym: O. campestris (L.) DC. var. johannensis Fern. (1899), Rhodora 1: 88].

T Canada: Newfoundland, St. John Island, 31.07.1925, leg. M.L. Fernald et al. 28615 (GH) holotype.

2n= 48 (6x).

2nD Hedberg (1967).

G CAN


Comments:

(1) Seen as a variety of O. campestris by Barneby (1952) and for priority reasons as O. campestris var. minor by Gillett et al. (1999 onwards). If this entity is considered part of O. campestris s. lat., it is a major geographical race and should be named as a subspecies (rather than as a variety) according to our rules. No subspecific combination is cited in the synonymy of Gillett et al. (1999 onwards). Moved here to the O. campestris aggregate from a position near O. arctica in Yurtsev's draft. (Elven)



(2) Oxytropis johannensis is proposed as a separate species in Yurtsev's draft but is fully synonymised with the 'terrae-novae' entity by almost all more recent North American authors. I have therefore reduced it to synonymy. It should probably be typified on a collection from near Quebec City (see Scoggan 1978). (Elven)

WARNING! Should probably be recombined as a subspecies of O. campestris.


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