Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 135




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Sect. Baicalia (Steller) Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 135.

B Baicalia Steller ***

Comments:

(1) All the four species of this 'semi-verticillate' section included by Fl. Arct. URSS (O. uniflora was described later) are local and with one exception known only from a few sites: O. putoranica from Taimyr - Putorana (3 sites mapped), O. sverdrupii from Ayon Island only (2 sites mapped), O. wrangelii from Wrangel Island (frequent) and E Chukotka (2 sites mapped), and O. schmorgunoviae from W Chukotka and further south (5-6 sites mapped). From this pattern, it is difficult to avoid the impression of local differentiation within a more major entity, even if several ploidy levels are known. The entities need some more justification before full acceptance.

The two North American species included in this group are treated very differently by American authors. Oxytropis bellii is considered by Gillett et al. (1999 onwards) as a variety of O. arctica. These authors do not assign the same importance to semiverticillate leaflets as does Yurtsev. Oxytropis splendens is not included by Gillett et al. as it does not reach the Arctic Archipelago but is accepted as a species by Porsild & Cody (1980) and Cody (1996). (Elven)
60.6.32  Oxytropis schmorgunoviae Jurtz. (1978), Byull. Mosk. Obshch. Ispyt. Prir., Biol. 83, 1: 125.

S

2n= 48 (6x).



2nD ?

G RFE


Comments:

(1) Var. tundrarum Jurtz. (1978), Byull. Mosk. Obshch. Ispyt. Prir., Biol. 83, 1: 126. In the type locality of var. schmorgunoviae: the upper reaches of Pineiveem River, at the edge of the right bank high terrace, but in the dry tundra surroundings of relic steppe islets. (Yurtsev)


60.6.33  Oxytropis sverdrupii Lynge (1929), Norw. N. Polar Exp. "Maud" Sci. Res. 5, 1: 8.

S

T [Described from NE Asia: Ayon I. Type in O.]



2n= 48 (6x).

2nD Zhukova & Tikhonova (1971).

G RFE

Comments:



(1) An extremely local species (Ayon Island), if it has not been found in more sites lately. (Elven)
60.6.34  Oxytropis wrangelii Jurtz. (1978), Byull. Mosk. Obshch. Isp. Prir., Biol. 83, 1: 127.

S

2n= 64 (8x).



2nD Zhukova (1966).

G RFE


Comments:
60.6.35  Oxytropis splendens Dougl. ex Hook. (1840), Fl. Bor.-Amer. 1: 147.

S

T [Described from: "Red River and south towards Pembina"]



2n=

2nD


G CAN

Comments:


60.6.36  Oxytropis bellii (Britt. ex Macoun) Palib. (1910), Bull. Soc. Bot. Geneve, ser. 2, 2: 19.

B Spiesia belli Britt. ex Macoun (1894), Canad. Record Sci. 1894: 1148 [?].

S O. arctica R. Br. var. bellii (Britt. ex Macoun) Boivin ***

2n=


2nD

G CAN


Comments:

(1) Included as a variety of O. arctica by Gillett et al. (1999). These authors do not attach the same importance to semiverticillate versus opposite leaflets as Yurtsev does. (Elven)


60.6.37  Oxytropis putoranica M. Ivanova (1976), Fl. Putorana 75.

S O. norinii Yu. Kozhevn. (1981), Bot. Zhurn. 66, 12: 1735.

2n= 16 (2x).

2nD ?


G SIB

Comments:

(1) In the type population near Baselak Lake in the Eastern Putorana Plateau. 2n=16 (Polozhij 1994). (Yurtsev)
60.6.38  Oxytropis uniflora Jurtz. (1988), Bot. Zhurn. 73, 12: 1650.

S

2n= 16 (2x).



2nD ?

G RFE


Comments:
Sect. Gloeocephala Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 108.

Comments:

(1) The North American representatives of this 'sticky' section have recently mostly been united within one species   O. borealis DC.   partly only as synonyms (Cody 1996), partly as intergrading varieties of the American type, i.e, probably subspecies in our concept (Welsh 1990, Gillett et al. 1999 onwards). Scoggan (1978) unite them within O. leucantha (see above), here considered in another section. We should consider wider species with subspecies as an alternative to the fairly numerous species of Yurtsev's draft and also see what can be done with the Russian representatives. The oldest species name in the aggregate seems to be O. borealis DC. 1825. (Elven)
60.6.39  Oxytropis vasskovskyi Jurtz. (1986), Fl. Arct. URSS 9, 2: 143, 181.

S

T Russian Far East: In cursu superiore fl. Kolymae, ad fl. Bochapczá, 1952, leg. A.P. Vasskovsky (LE) holotype.



2n= (1) 16 (2x). (2) 32 (4x).

2nD ?


G RFE

Comments:

(1) A fairly wide-ranging entity from Kharaulakh to Anjui Mts. (Elven)
60.6.40  Oxytropis inopinata Jurtz. (1986), Fl. Arct. URSS 9, 2: 144, 182.

S

T Siberia: Jacutia, ad Lenam inferiorem, in glareis inundatis fl. Lenae prope ostium fl. Neleger, 16.07.1984, leg. V.V. Petrovsky (LE) holotype.



2n= 16 (2x).

2nD ?


G SIB

Comments:

(1) A very local entity in the lowermost reaches of Lena, not very distant from the nearest (isolated) site of O. vasskovskyi. Its relationship to that species should be critically evaluated, as a possible marginal population group. (Elven)

WARNING! In danger of merge with O. vasskovskyi.
60.6.41  Oxytropis kateninii Jurtz. (1986), Arkt. Fl. SSSR 9,2: 146, 182.

S

T Russian Far East: Peninsulae Czukotskij (Czukotensis) pars boreali-orientalis, regio ripae dextrae fl. Chesmymken adjacens, 2 km ab ostio, 08.08.1982, leg. A.E. Katenin (LE) holotype.



2n=

2nD


G RFE

Comments:

(1) Reported from a single site in Chukchi Peninsula, within the range there of O. borealis. Its relationship to the latter must be analysed before acceptance. (Elven)

WARNING! In danger of merge with O. borealis.
60.6.42  Oxytropis viscida Nutt. ex Torr. & Gray (1838), Fl. N. Amer. 1: 341.

S O. borealis DC. var. viscida (Nutt. ex Torr. & Gray) ***

T [Described from: Rocky Mountains, near the source of the Oregon]

2n= (1) 16 (2x). (2) 32 (4x).

2nD ?

G ALA


Comments:

(1) Chromosome counts for non-arctic populations. (Yurtsev)

(2) If we consider more widely delimited species, this will be a Cordilleran subspecies of O. borealis. (Elven)

WARNING! Should probably be reduced to a subspecies of O. borealis.
60.6.43  Oxytropis borealis DC. (1825), Prodr. 2: 275.

S O. borealis DC. var. borealis sensu Welsh (1990) and Gillett et al. (1999); O. leucantha auct., non (Pall.) Bunge (1874).

T Russian Far East: "e sinu S. Laurentii in terra Tschuktschorum septentrionem versus a fretus Beringii, 1825, leg. Fischer (G-DC) holotype.

2n= 48 (6x).

2nD Holmen (1962); Johnson & Packer (1968 Ala, 2n=c. 48).

G RFE ALA

Comments:

(1) In CH.11 a non typical form. (Yurtsev)

(2) The demarcation with the following taxon should be specified. (Yurtsev)

(3) Acc. to Welsh (1990), O. borealis is seen as a wide species with several intergrading varieties (varieties in the American concept, probably subspecies in the European-Russian concept). At least O. borealis s. str., O. glutinosa, O. hudsonica and O. viscida fall inside this extended O. borealis. There might, however, be a ploidy problem. (Elven)

(4) This entity could be merged with O. glutinosa as a more widely delimited O. borealis subsp. borealis, or the two could be kept apart as two subspecies. As two species, both present on the North American side of the Bering Strait, they are much too weakly characterised. (Elven)
60.6.44  Oxytropis glutinosa A.E. Porsild (1951), Bull. Natl. Mus. Canada 121: 240.

S O. borealis DC. var. viscida (Nutt. ex Torr. & A. Gray) *** p.p.

2n= 16 (2x).

2nD Mulligan & Porsild in Löve (1970b).

G ALA CAN

Comments: See O. borealis above.

(1) The pattern in N America might justify four regional races: 'borealis' as amphi-Beringian and Alaskan, 'glutinosa' as NW Canadian, 'viscida' as Cordilleran, and 'hudsonica' as C Canadian ('Hudsonian'). An alternative would be to merge 'borealis' and 'glutinosa' within one subspecies. (Elven)

WARNING! Should either be reduced to a separate subspecies of O. borealis or merged with a subsp. borealis.
60.6.45  Oxytropis hudsonica (E. Greene) Fern. (1928), Rhodora 30: 142.

B Aragallus hudsonicus E. Greene (1905), Proc. Biol. Soc. Wash. 18: 17.

S O. viscida Nutt. subsp. hudsonica (E. Greene) Á. Löve & D. Löve (1982), Taxon 31: 347; O. borealis DC. var. hudsonica (E. Greene) S.L. Welsh (1991), Great Basin Natural. 50: 357.

T Canada: Whale River, Hudson Bay, 24.06.1896, leg. A.P. Low 14272 (NDG) holotype.

2n=

2nD


G CAN

Comments:

(1) A treatment as subspecies is in accordance with Gillett et al. (1999 onwards, as variety) considered their definition of variety. (Elven)

WARNING! Should probably be reduced to a subspecies of O. borealis.
60.6.46  Oxytropis middendorffii Trautv. (1847), Fl. Taim. 49.

S

Comments:



(1) The splitting of this Siberian species into seven arctic subspecies seems excessive but it is difficult to evaluate it as long as the descriptions are not translated. There is also an eight subspecies (subsp. jarovii) that approaches the Arctic between Lena and Kolyma. The proposed subspecies are mainly allopatric to parapatric but with comparatively narrow ranges: 'middendorffii' from Taimyr to Lena, 'schmidtii' in one site just south of the preceding in SW Taimyr, orulganica' in Kharaulakh, ('jarovii' between Kharaulakh and Kolyma), 'coerulescens' mainly east of Kolyma in E Chukotka but isolated in the Koryak Mts, 'anadyrensis'mainly in Chukotka but isolated much farther west, 'submiddendorffii' disjunct in Wrangel Island and Chukotka, and 'trautvetteri' acc. to Fl. Arct. URSS far south of the Arctic along the Okhotsk Sea.

These entities must be well justified before we can accept them as major geographic races, i.e., subspecies. (Elven)



WARNING! All subspecies might be omitted if not well justified.
60.6.46.1  Oxytropis middendorffii Trautv. subsp. middendorffii

S

2n= 48 (6x).



2nD Löve & Löve (1975) list five Russian counts, all as arctic.

G SIB


Comments:
60.6.46.2  Oxytropis middendorffii Trautv. subsp. orulganica Jurtz. (1968), Fl. Suntar-Khayata 72.

S

2n= 48 (6x).



2nD ?

G SIB


Comments:
60.6.46.3  Oxytropis middendorffii Trautv. subsp. submiddendorffii Jurtz. (1986), Fl. Arct. URSS 9, 2: 137, 181.

S

T Russian Far East: Insula Wrangelii, prope ostium rivuli Chrustalnyi [affluentiae sinistrae fl. Mamontovajae in curso medio], 28.07.1965, leg. P.G. Zhukova & V.V. Petrovsky (LE) holotype.



2n= 48 (6x).

2nD ?


G RFE

Comments:


60.6.46.4  Oxytropis middendorffii Trautv. subsp. coerulescens Jurtz. & Petrovsky in Jurtz. (1986), Fl. Arct. URSS 9, 2: 137, 181.

S

T Russian Far East: Terra Tschuktschorum occidentalis, planities montium Anjunensis, systema fl. Pogynden, fl. Alarmagtyn in cursu medio, rivul. Jagodnyi, 06.07.1974, leg. V.V. Petrovsky & T.M. Koroleva (LE) holotype.



2n= 48 (6x).

2nD ?


G RFE

Comments:


60.6.46.5  Oxytropis middendorffii Trautv. subsp. schmidtii (Meinsh.) Jurtz. (1968), Fl. Suntar-Khayata 74.

B O. schmidtii Meinsh. in Schmidt (1872), Fl. Jeniss. 97.

S

2n= 48 (6x).



2nD ?

G SIB


Comments:

(1) This entity is especially suspect! (Elven)


60.6.46.6  Oxytropis middendorffii Trautv. subsp. anadyrensis (V. Vass.) Jurtz. (1980), Byull. Mosk. Obshch. Isp. Prir., Otd. Biol. 85, 6: 104.

B O. anadyrensis V. Vassil. (1960), Bot. Mater. Gerb. Bot. Inst. AN SSSR 20: 248.

S

2n= 48 (6x).



2nD ?

G RFE


Comments:
60.6.46.7  Oxytropis middendorffii Trautv. subsp. trautvetteri (Meinsh.) Jurtz. (1986), Fl. Arct. URSS 9, 2: 141.

B O. trautvetteri Meinsh. in Schmidt (1872), Fl. Jeniss. 97.

S

2n=


2nD

G RFE


Comments:

(1) Gursenkov & Pavlova (1984) rcorded 2n=16 for the specimens from the coast of Sea of Okhotsk, which looks unexpected due to a significant morphological similarity of the Okhotsk race with 2n=48. The above records are to be confirmed on fresh, greater material. The specimens from CH.11 (Pekuluei Range) are not typical and are, in fact, intermediate to subsp. anadyrensis. (Yurtsev)



(2) Therefore, I think we at least should omit this subspecies and reduce it to a comment under subsp. anadyrensis if subspecies are retained. (Elven)


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