Annotated checklist of the weevils (Curculionidae sensu lato ) of North America, Central America, and the West Indies

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Annotated checklist of the weevils (Curculionidae sensu lato )

of North America, Central America, and the West Indies

(Coleoptera: Curculionoidea)
Charles W. O'Brien and Guillermo J. Wibmer
This checklist treats the names of the 843 genera and 7,068 species (as well as their synonyms) currently recognized as valid that are found in the New World north of South America (except for those from Trinidad and Tobago which will be considered in a subsequent publication on South American Curculionidae sensu lato).
The idea for a weevil checklist originated with Ross Arnett, Jr. as part of the North American Beetle Fauna Project. When this project was terminated in 1980, we decided to expand the scope of the checklist and publish it in an annotated form. While it owes its origin to the NABFP it is published independently of that organization.
The checklists of the weevils of North America (Leng 1920, Leng and Mutchler 1927 and 1933, Blackwelder 1939, and Blackwelder and Blackwelder 1948), and of Mexico, Central America, the West Indies and South America (Blackwelder 1947) have become increasingly outdated because of numerous revisions and descriptions of new taxa. In this list we have added many new distribution records as well.
We have attempted to follow the current classifications of most specialists, as published. For this reason the classification used here is not identical with that of the Coleopterorum Catalogus, the two checklists mentioned above, nor that used by Kissinger (1964).
We have tried to verify all citations by checking original references and those "Not seen" are so marked in the bibliography. In the process of checking previous catalogs, checklists, and other early references, we have located many homonymies and incorrect synonymies that require name changes if we are to remain in accord with the International Code of Zoological Nomenclature. It is unfortunate that this will require changing the names of many well known genera, some that include important economic species, but we feel that in these cases the use of strict priority must be followed to avoid potential chaos in nomenclature. Most of these synonymies have been known by specialists for years, some for more than 150 years, and they have been repeated incorrectly in catalogs, checklists, and even monographs. It is clear that many workers have not considered as available any name from the lists of Germar (1817) or Dejean (1821, 1833--1837, and 1837), but all genus-group names in these publications must be carefully checked to determine which are available. Those genus-group names that include any available species-group names must be treated as available, in spite of the lack of descriptions. Schoenherr himself did not consider his generic names listed in 1823 and 1825 as available from those dates, but many of these are. Because many of the name changes caused by the strict application of the Law of Priority will upset long established names, we suggest any specialist who wishes to do so may apply to the International Commission for conservation of the more recent names and rejection of the older names. We checked all volumes of Neave (1939-1940, 1950), Edwards and Hopwood (1966), and Edwards and Vevers (1975) for homonymies, and discovered a number that have been overlooked by taxonomists. Below is a list of homonymies (with new names), as well as other lists which we believe will benefit indexers and weevil taxonomists.

2Mem. Amer. Ent. Inst., no. 34, 1982

The bibliography lists all references cited in addition to many other selected references. We have included a list of abbreviations, with complete citations of abbreviated periodicals. With a very few exceptions we follow the "NCPTWA (National Clearinghouse for Periodical Title Word Abbreviations) Word-Abbreviation List", 1971 edition.
The keys and revisions cited in the checklist have been selected from those known to us. We have not included all keys, but only those that we considered significant. When there is a key or revision to a group that has been split, the key or revision is cited under each name. We did not cite the keys in LeConte and Horn (1876), Blatchley and Leng (1916), Kissinger (1964), nor Hutch (1971), and these publications should be checked for keys, especially if other keys are not cited. The keys in LeConte and Horn (1876) tend to be incomplete because of the many additions to the known fauna since that time. Blatchley and Leng (1916), and Hutch (1971) key subfamilies, genera and species on a regional basis, the eastern United States and eastern Canada in the former, and the Pacific Northwest in the latter. Kissinger (1964) has keys to most North American subfamilies and most genera. O'Brien and Wibmer (1981) provide an annotated bibliography of keys to Latin American weevils.
We have included all spelling errors known to us of all genus-group names. We list the first such error that we know. We have not included all spelling errors of species -group names, but have treated these selectively, primarily including those that we believe might cause confusion. We have attempted to correct the endings of species -group names to conform with the gender of their current genus. However, synonyms are listed with their original spelling.
Synonyms of introduced species are listed only when they have been used in New World literature. We have included a few synonyms based on Old World genera or species which have been reported since the last catalogs on the weevils (e.g., Coleopterorum Catalogus (1934, 1936, 1937), Leng Catalog Supplement 5 (Blackwelder and Blackwelder 1948), and the Blackwelder checklist, part 5 (1947)).
Even though varieties and forms have no status in Zoological Nomenclature, we are including them as such, unless they have been synonymized, or their status changed. We prefer not to lose the information on the distribution associated with these names. It is possible that some of them may be treated in the future as valid subspecies or species.
Nomina nuda are cited for names included in previous lists or synonymies to make it clear why such names are not available.
The page of publication (following the year of publication) is the page on which the species description begins, not the page of use in a key, unless the name occurs only in the key, for which we then say "[in key]". If the name was published at an earlier date in a key and the description published later, the page for the description is cited also, in brackets.
An * is used to indicate that a genus and/or species has been introduced into the region covered by the checklist.
We have included in the checklist a few recently described fossil weevils, but have not included the genera nor species from Scudder (1893) and Wickham (1912), as well as from many small papers. See Wickham (1920) for a complete list of fossil Coleoptera including weevils described to that date.
We have used the 5 regions for North America outlined by Blackwelder et al. (1977), and we have added 2 regions.

Region (1) Northeastern United States and adjacent Canada: CT - Connecticut; DC - District of Columbia; DE - Delaware; GR - Greenland; Hud. Bay T. - Hudson Bay
Territory; IL - Illinois; IN - Indiana; LB - Labrador; MA - Massachusetts; MD - Maryland; ME - Maine; MI - Michigan; NB - New Brunswick; NE - Newfoundland; NH - New Hampshire; NJ - New Jersey; NS - Nova Scotia; NY - New York; OH - Ohio; ON - Ontario; PA - Pennsylvania; PE - Prince Edward Island; PQ - Quebec; RI - Rhode Island; SP - Saint Pierre & Miquelon; VT - Vermont; WI - Wisconsin; L. Sup. - Lake Superior Region; N. Eng. - New England.

Region (2) Southeastern United States: AL - Alabama; AR- Arkansas; FL -

Florida; GA - Georgia; KY - Kentucky; LA - Louisiana; MS- Mississippi; NC - North Carolina; SC - South Carolina; TN - Tennessee; VA - Virginia; WV - West Virginia.

Region (3) Southwestern United States, and Baja California, Mexico: AZ- Arizona;

BJ - Baja California; CA - California; NM - New Mexico; OK - Oklahoma; TX - Texas.

Region (4) Northwestern United States and adjacent Canada, including Alaska and

the Aleutian Islands: AK - Alaska; BC - British Columbia; ID - Idaho; NV - Nevada; OR - Oregon; UT- Utah; WA - Washington.

Region (5) North Central United States and adjacent Canada: AB - Alberta; CO - Colorado; IA - Iowa; KE - Keewatin; KS - Kansas; MB - Manitoba; MC - Mackenzie; MN- - Minnesota; MO - Missouri; MT - Montana; ND - North Dakota; NE- Nebraska; NT - -Northwest Territories; SA - Saskatchewan; SD - South Dakota; WY - Wyoming; YU - Yukon Territory.

Region (6) Mexico (excluding Baja California), and Central American countries:

Belize - (=British Honduras); C. Rica - Costa Rica; Cocos Is. - Cocos Island (Costa Rica); El Salv. - El Salvador; Guat. - Guatemala; Hond. - Honduras; Mex. - Mexico; Nic. - -Nicaragua; Pan. - Panama; C. Am. - Central America.

Region (7) West Indies (excluding Trinidad and Tobago): Dom. Rep. - Dominican Republic; I. Pinos - Isla de Pinos (Cuba); Jam. - Jamaica; P. Rico - Puerto Rico; W. Indies - West Indies; [other West Indian localities not abbreviated].

Miscellaneous Abbreviations: N. Am. - North America; S. Am. - South America;

U.S.A. - United States of America.
The distributions cited are based on previously published records (not including those found only in checklists) and on specimens in the senior author's collection. Doubtful records from early literature (presumably based on misidentifications) are not included. Various specialists have added and deleted some localities also.
New Names
Acalles hustachei O'Brien & Wibmer [=A. squamosus Hustache 1930 (not Solari & Solari 1907, Ann. Mus. Civ. Stor. Nat. Genova, ser. 3, 3:535)]

Acalles neoclathratus O'Brien & Wibmer [=A. clathratus Fiedler 1942 (S. Am.) (not LeConte 1876, Proc. Am. Philos. Soc. 15(96:242)]

Anthonomus neosolani O'Brien & Wibmer [=A. solani Ferragu 1963 (not Fall 1913, Trans. Am. Entomol. Soc. 39:49)]

Arthleptrus Burke (subgenus of Anthonomus) [=Leptarthrus Dietz 1891, type-species Anthonomus julichi Dietz 1891, by present designation, H. R. Burke (not Stephens 1829, Nomencl. British Insects, p. 55 [Diptera])]

Gilbertiola Osella [=Gilbertia Osella 1977, type-species Raymondionymus helferi Gilbert 1956, by original designation (not Cossmann 1889, Ann. Soc. R. Malacol. Belg. 24:347 [Mollusca]; not Jordan & Eigenmann 1890, Bull. U. S. Fish Comm., 1888(1890), 8:333, 346 [Pisces]; not Walsingham 1891, Entomol. Mon. Mag. 27:259 [Lepidoptera])]

Neocimberini O'Brien & Wibmer [=Rhinomacerini of authors; =Cimberini of authors]

Neocimberis O'Brien & Wibmer [type-species Rhinomacer attelaboides Fabricius 1787, by present designation; =Rhinomacer of authors (not Fabricius 1781, Species insectorum, p. 199 [Coleoptera]); =Cimberis of authors (not des Gozis 1881, Bull. Soc. Entomol. Fr., p. CXII [Coleoptera])]

Neodiplogrammus O'Brien & Wibmer [=Diplogrammus Chevrolat 1875, type-species Rhynchaenus quadrivittatus Olivier 1807, by present designation (not Gill 1865, Ann. Lyceum Nat. Hist. N. Y. 8:143 [Pisces])]
4Mem. Amer. Ent. Inst., no. 34, 1982
Neoerethistes O'Brien & Wibmer [=Erethistes Pascoe 1872, type-species Cholus tetricus Fahraeus 1844, by present designation (not Mueller & Troschel 1849, Horae Ichthyol. (3):12 [Pisces])]

Neoperitelinus O'Brien & Wibmer [=Peritelinus Solari 1955, type-species Peritelus angulicollis Desbrochers 1897, by monotypy (Old World) (not Casey 1888, Ann. N. Y. Acad. Sci. 4:263)]

Neophrynoides O'Brien & Wibmer [=Phrynoides Chevrolat 1885, type-species Phrynoides luteus Chevrolat 1885, by monotypy (not Simon 1864, Hist. Nat. Araig ees, p. 437 [Arachnida])]

Neophycocoetes O'Brien & Wibmer [=Phycocoetes LeConte 1876, type-species Phycocoetes testaceus LeConte 1876, by monotypy (not Agassiz 1846, Nomencl. zool., Index universalis, p. 288 [Pisces])]

Neoulosomus O'Brien & Wibmer [=Ulosomus Schoenherr 1826, type-species Ulosomus erinaceus Schoenherr 1826, by original designation (not Schoenherr 1825, Isis Oken, column 585 [Coleoptera])]

Neozurus O'Brien & Wibmer [=Zurus Heller 1895, type-species Zurus aurivillianus Heller 1895, by present designation (not Amyot 1846, Ann. Soc. Entomol. Pr., ser. 2, 4:162 [Hemiptera] )]
New Synonymies
Acalles Schoenherr 1825 [=Ulosomus Schoenherr 18251

Acalles subhispidus LeConte 1878 [=Pseudoacalles maculatus Blatchley 1920]

Apteromechus nitidifrons Champion 1906 [=A. rugipectus Champion 1906]

Apteromechus pumilus (Boheman 1837) [=A. microstictus Fall 1925]

Bagous nebulosus LeConte 1876 [=B. ochraceus Blatchley 1916]

Bagous transversus LeConte 1876 [=B. carinatus Blatchley 1925]

Calandrinus grandicollis LeConte 1876 [=C. angustulus Casey 1920]

Calandrinus obsoletus Casey 1892 [=C. insignis Casey 1892]

Cleonis canescens (LeConte 1876) [-Cleonus grandirostris Casey 1891]

Cossonus deceptor Boheman 1838 [=C. nitidicollis Champion 1909]

Cryptolepidus Van Dyke 1936 [=Pseudoeucyllus Tanner 1950]

Cryptorhynchus duplaris (Champion 1906) [=Cryptorrhynchus subcylindricus Champion 1906]

Cylindrocopturus operculatus (Say 1824) [=Copturodes missourianus Casey 1897]

Dirabius rectirostris (LeConte 1876) [=Limnobaris tenua Blatchley 1916]

Epicaerus Schoenherr 1834 [=Cacochromus Sharp 1891; =Epagrius Schoenherr 1840]

Eudiagogus pulcher Fahraeus 1840 [=E. pogo Warner 1979]

Geoderces puncticollis Casey 1888 [=Nemocestes sordidus Van Dyke 1936]

Gerstaeckeria Champion 1905 [=Opuntiaphila Pierce 1912; =Philopuntia Pierce 1912]

Listronotus delumbis (Gyllenhal 1834) [=Anchodemus schwarzi LeConte 1876; =Listroderes solutus Boheman 1842; =Listroderes spurcus Boheman 1842; =Macrops indistinctus Dietz 1889]

Listronotus echinatus (Dietz 1889) [=Macrops schauppii Angell 1893]

Listronotus hornii (Dietz 1889) [=Macrops setiger Dietz 1889]

Listronotus humilis (Gyllenhal 1834) [=Macrops subcribratus Dietz 1889]

Listronotus hyperodes (Dietz 1889) [=Macrops nevadensis Dietz 1889]

Listronotus maculicollis (Kirby 1837) [=Macrops mirabilis Dietz 1889; =M. myasellus Dietz 1889]

Listronotus porcellus (Say 1831) [=Hyperodes minimus Blatchley 1916]

Listronotus sparsus (Say 1831) [=Listroderes immundus Boheman 1842; =Listroderes latiusculus Boheman 1842; =Macrops imbellis Dietz 1889; =M. Iongulus Dietz 1889]

Listronotus vitticollis (Kirby 1837) [=Macrops interpunctatulus Dietz 1889; =M. interstitialis Dietz 1889; =M. tenebrosus Dietz 1889]

Lixellus lutulentus (Boheman 1843) [=Anchodemus angustus LeConte 1876]

Metamasius aurocinctus (Champion 1910) [=Eucactophagus albipunctata Champion 1910; =E. albopunctatus Csiki 1936]


Nedyus Schoenherr 1825 [=Cidnorhinus Thomson 18591

Neocimberis pilosus (LeConte 1876) [=Rhinomacer pallipennis Blatchley 1916]

Nicentrus grossulus Casey 1893 [=N. parallelus Casey 1920]

Ophryastes geminatus (Horn 1876) [=O. vittatus Hatch 1971 (not Say 1824)]

Pantomorus femoratus Sharp 1891 [=P. robustus Sharp 1891]

Pentarthrum elumbe (Boheman 1838) [=P. cylindricum Wollaston 1861; =P. Wollaston 1873]

Phyllotrox sejunctus (Fall 1913) [=Euclyptus equisectus Fall 1913]

Phyrdenus setiferus (Boheman 1844) [=P. caseyi Faust 1896]

Phytobius Schoenherr 1833 [=Litodactylus Redtenbacher 1845 (or 1849)]

Rhinoncus asperulus (Dietz 1896) [=Mecopeltus aeneosquamosus Dietz 1896]

Rhinoncus pusillus (Dietz 1896) [=Mecopeltus scandens Dietz 1896]

Rhinoncus squamosus (LeConte 1876) [=Pelenomus impressiventris Blatchley 1916]

Rhynchites Schneider 1791 [=Platyrynchus Thunberg 1815]

Rhyssomatus strangulatus Gyllenhal 1837 [=R. barioides Fiedler 1937]

Thecesternus affinis (LeConte 1857) [=Lithodus morbillosus LeConte 1857; =L. rectus LeConte 1857]

Udeus Champion 1902 [=Moracetribus Bondar 1957]

Xystus Schoenherr 1826 [=Stegotes Casey 1922]
Resurrected Names
Alceis Billberg 1820 [not a nomen nudum; senior synonym of Naupactus Dejean 1821 and Iphius Schoenherr 1823]

Ampeloglypter ampelopsis (Riley 1869) [not a nomen nudum; senior synonym of A. ater LeConte 1876]

Anthonomus rutilus (Boheman 1843) [senior synonym of A. nubiloides Fall 1928]

Apteromechus pumilus (Boheman 1837) [not a synonym of Eubulus bisignatus (Say 1831)]

Archarias Dejean 1821 [not a nomen nudum; senior synonym of Aphyoramphus Guérin 1844 (and has priority over the subgenus Cholus Germar 1824)]

Campylirhynchus Dejean 1821 [not a nomen nudum ]

Ceutorhynchus squamosulus Sleeper 1957 [not a homonym of C. squamulosus Brisout 1869]

Cleonis Dejean 1821 [not a nomen nudum; senior synonym of Geomorus Schoenherr 1823 and Cleonus Schoenherr 1826]

Cleonis canescens (LeConte 1876) (Cleonus) [not a homonym of Lixus canescens (Chevrolat 1873) (Xanthochelus)]

Cratosomus curassaviensis (Voet 1806) [not a nomen nudum; senior synonym of C. stellio (Olivier 1807)]

Cryptorhynchus sulcicollis Suffrian 1876 [not a synonym of C. subnotatus Boheman 1837]

Curculio obtusus (Blanchard 1884) (Balaninus) [not a homonym of Tropiphorus obtusus (Bonsdorff 1785) (Curculio )]

Curculio uniformis (LeConte 1857) (Balaninus) [not a homonym of Phyllobius uniformis (Marsham 1802)(Curculio)]

Diaprepes marginatus (Fabricius 1775) [senior synonym of D. bivittatus (Fabricius 1787)]

Dorytomus leucophyllus (Motschulsky 1845) [senior synonym of D. subsignatus (Mannerheim 1853)]

Eccoptus Dejean 1821 [not a nomen nudum; senior synonym of Zygops Schoenherr 1825]

Eugnamptus angustatus (Herbst 1797) [senior synonym of E. collaris (Fabricius 1801)]

Geoderces Horn 1867 [incorrectly replaced with Nemocestes Van Dyke 1936]

Gerstaeckeria cactophaga Pierce 1912 [not a synonym of G. basalis (LeConte 1876)]

Graptus Schoenherr 1823 [not a nomen nudum; senior synonym of Alophus Schoenherr 1826) (Old World)]

Hilipimorphus Champion 1905 [not a homonym of Hilipomorphus Desbrochers 1891]

Himatium Wollaston 1873 [not a homonym of Imatium Clark 1860; senior homonym of Himatium Gemminger & Harold 1876]

6Mem. Amer. Ent. Inst., no. 34, 1982

Laemosaccus nephele (Herbst 1797) [senior synonym of L. plagiatus (Fabricius 1798)]

Lepesoma Motschulsky 1845 [senior synonym of Dyslobus LeConte 1869]

Litostylus diadema (Fabricius 1787) [senior synonym of L. juvencus (Olivier 1807)]

Maemactes subfasciatus Champion 1905 [not a synonym of M. perforatus Champion 1905)]

Menoetius Dejean 1821 [not a nomen nudum; senior synonym of Ptilopus Schoenherr 1823 and Lachnopus Schoenherr 1840]

Nanodes Schoenherr 1825 [not a nomen nudum; senior homonym of Nanodes Stephens 1826; senior synonym of Nanophyes Schoenherr 1838]

Nedyus Schoenherr 1825 [not a nomen nudum; senior synonym of Cidnorhinus Thomson 1859]

Notiodes Schoenherr 1838 [senior synonym of Endalus Laporte 1840]

Onychylis guadelupensis (Hustache 1929) [not a synonym of O. setiger Champion 1902]

Oxyderces Schoenherr 1823 [not a synonym of Compsus Schoenherr 1823; senior synonym of Plococompsus Marshall 1922]

Pachyrhinus Schoenherr 1823 [not a nomen nudum ; senior synonym of Scythropus Schoenherr 1826] Peridinetus roeselii (Boheman 1836) [senior synonym of P. maculatus Rosenschoeld 1837]

Toxophorus Dejean 1835 [not a nomen nudum; not a homonym of Toxophora Meigen 1803; senior synonym and senior homonym of Toxophorus Schoenherr 1836]

Tropirhinus Schoenherr 1823 [not a nomen nudum ; senior synonym of Callizonus Schoenherr 1826]

Xystus Schoenherr 1826 [not a nomen nudum; senior synonym of Stegotes Casey 1922]
New Combinations
Acalles maculatus (Blatchley 1920) [from Pseudoacalles; new synonym of A. subhispidus LeConte 1878] Apteromechus pumilus (Boheman 1837) [from Eubulus]

Archarias, all species [from Cholus]

Bagous belfragei (Burke 1963) [from Pnigodes]

Bagous buchanani (Tanner 1943) [from Pnigodes]

Bagous tuberosus (Tanner 1943) [from Pnigodes]

Campylirhynchus, all species [from Rhinoncus]

Chalcodermus pupillatus (Suffrian 1872) [from Rhyssomatus]

Championius latescens (Champion 1905) [from Acalles]

Championius rugirostris (Champion 1905) [from Acalles]

Cimbocer apilosa (Sharp 1891) [from Amphidees]

Cleonis, all species [from Cleonus]

Cophes yucatanus (Champion 1906) [from Cryptorhynchus]

Cryptolepidus boulderensis (Tanner 1950) [from Pseudoeucllus]

Dirabius tenua (Blatchley 1916) [from Limnobaris ; new synonym of D. rectirostris (LeConte 1876)]

Eccoptus, all species [from Zygops]

Epicaerus brevirostris (Sharp 1891) [from Epagrius]

Epicaerus metallescens (Boheman 1840) [from Epagrius; synonym of E. nubilosus (Boheman 1840)]

Epicaerus nubilosus (Boheman 1840) [from Epagrius ]

Epicaerus retrorsus (Champion 1911) [from Epagrius]

Epicaerus tolucae (Sharp 1891) [from Epagrius; synonym of E. nubilosus (Boheman 1840)]

Eucyllus nevadensis (Casey 1888) [from Thinoxenus ]

Eurhoptus binodulus (Champion 1910) [from Acalles ]

Eurhoptus tantillus (Champion 1905) [from Acalles ]

Geoderces, all species [from Nemocestes]

Gilbertiola helferi (Gilbert 1956) [from Gilbertia]

Gilbertiola schusteri (Gilbert 1956) [from Gilbertia]


Hilipimorphus forreri Champion 1905 [from Hilipomimus]

Himatium, all species [from Himatinum]

Listronotus nigropunctatus (Suffrian 1870) [from Listroderes]

Maemactes hispidulus (LeConte 1876) [from Sternoeoelus]

Menoetius, all Species, exeept M. luctuosus (Gyllenhal) and M. Iuctuosus (Klug) [from Lachnopus]

Menoetius luctuosus (Gyllenhal 1834) [from Exophthalmus; synonym of M. luctuosus (Klug 1829)]

Menoetius luctuosus (Klug 1829) [from Exophthalmus]

Micromimus corticalis (Boheman 1845) [from Caulophilus]

Micromimus politus (Casey 1892) [from Caulophilus; synonym of M. corticalis (Boheman 1845)]

Nanodes, all species [from Nanophyes]

Nedyus, all species [from Cidnorhinus]

Neocimberis, all species [from Cimberis]

Neodiplogrammus, all species [from Diplogrammus]

Neoerethistes, all species [from Erethistes]

Neoperitelinus angulicollis (Desbrochers 1897) [from Peritelinus; Old World]

Neophrynoides luteus (Chevrolat 1885) [from Phrynoides]

Neophycocoetes testaceus (LeConte 1876) [from Phyeocoetes]

Neoulosomus, all species Ifrom Ulosomus]

Neozurus, all species [from Zurus]

Notiodes, all species [from Endalus]

Odontocorynus falsus (LeConte 1876) [from Centrinaspis]

Oxyderces, all species [from Plococompsus]

Pachyrhinus, all species [from Scythropus]

Pentarthrum elumbe (Boheman 1838) [from Rhyncolus ]

Pentarthrum fusiformis (Wollaston 1873) [from Rhyncolus; synonym of P. elumbe (Boheman 1838)]

Phytobius, all species [from Litodactylus]

Rhinoncus, all species [from Phytobius]

Sternocoelus erubescens (Champion 1906) [from Cryptorhynchus]

Toxophorus, all species [from Atenistes and Ludovix]

Tropirhinus, all species [from Callizonus]

Tyloderma pilosellum (Chevrolat 1879) Ifrom Euscepes]

Tyloderma pilosellum v. parvulum (Hustache 1930) [from Euscepes]

Tylodinus anchonoides (Champion 1910) [from Acalles]

Tylodinus cavicollis (ChHmpion 1905) [from Acalles]

Tylodinus cinericius (Champion 1905) [from Acalles]

Tylodinus indutus (Champion 1905) [from Acalles ]

Tylodinus quadrituberculatus (Champion 1905) [from Acalles]

Tylodinus rufotuberculatus (Champion 1905) [from Acalles]

Tylodinus rugulosus (Champion 1905) [from Acalles ]

Tylodinus triseriatus (Champion 1910) [from Acalles]

Tylodinus truquii (Champion 1910) [from Acalles ]

Udeus cecropiae (Bondar 1957) [from Moracetribus; S. Am.]

Udeus sergioi (Bondar 1957) [from Moracetribus ; S. Am.]

Xenosomina gonodera (Chevrolat 1879) [from Xenosomus]

Xenosomina inflata (Champion 1905) [from Gerstaeckeria]

Xenosomina turpis (Champion 1905) [from Gerstaeckeria]

Xystus, all species [from Stegotes]
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