5 Ophioglossaceae (Draft: N. N. Tzvelev) 1 Botrychium




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5  Ophioglossaceae (Draft: N.N. Tzvelev)

5.1  Botrychium Sw. (1801), J. Bot. (Schrader) 1800(2): 110.

Comments:

(1) In his draft, Tzvelev includes for Greenland also Botrychium pumicola Coville in Underwood (1900), Native Ferns, ed. 6, 69. Are there any indications that this narrow Oregon endemic occurs in Greenland? Possibly a misinterpretation of B. simplex. I have omitted it. Another species has been tentatively added (B. pinnatum).

The sequence of species has been changed from Tzvelev's draft, to keep together B. lanceolatum, B. boreale, and B. pinnatum, and B. lunaria and B. minganense.

Fl. N. Amer. 2 (1993) separates the eight species on three subgenera: Osmundopteris (J. Milde) R.T. Clausen (B. virginianum), Sceptridium (Lyon) R.T. Clausen (B. multifidum) and Botrychium (the others). If we accept subgenera in Equisetum, we should also consider inclusion of these.

The (very) narrow species concept of American pteridologists must be discussed as it here might result in a finer differentiation in species on the American than on the European -Asian side. (Elven)


5.1.1  Botrychium virginianum (L.) Sw. (1801), J. Bot. (Schrader) 1800, 2: 111.

B Osmunda virginiana L. (1753), Sp. Pl. 1064.

S

T Linnaean Herbarium 1244.3 (LINN) lectotype, selected by Proctor (1985, Ferns of Jamaica 51).



2n= 184 (4x).

2nD Fl. N. Amer. 2 (1993).

G CAN?

Comments:



(1) The plants that reach or nearly reach the Arctic are often (Hultén 1968, Porsild & Cody 1980) placed in a subsp. europaeum (Ångstr.) Clausen (1938), Mem. Torrey Bot. Club 19: 101 [basionym: B. virginianum (L.) Sw. var. europaeum Ångstr. (1854): Bot. Not. 1854: 68]. If we accept a subspecific split, we have to include an entry for the subspecies. Another question is whether the plant reaches the Arctic as defined. Scoggan (1978) gives the northernmost occurrences in the Hudson Bay region as Gillam (Manitoba) and Severn River (Ontario), 5545'N. (Elven)

WARNING! Will be excluded if arctic occurrence in Canada not confirmed.
5.1.2  Botrychium multifidum (S.G. Gmel.) Rupr. (1859), Beitr. Pflanzenk. Russ. Reiches 11: 40.

B Osmunda multifida S.G. Gmel. (1768), Novi Comment. Acad. Sci. Imp. Petrop. 12: 517.

S B. multifidum (S.G. Gmel.) Rupr. var. multifidum.

T [Described from Russia.]

2n= 90 (2x).

2nD Löve & Löve (1975) list three non-arctic counts; Fl. N. Amer. 2 (1993).

G CAN GRL

Comments:


5.1.3  Botrychium lanceolatum (S.G. Gmel.) Ångstr. (1854), Bot. Not. 1854: 68.

B Osmunda lanceolata S.G. Gmel. (1768), Novi Comment. Acad. Sci. Imp. Petrop. 12: 516.

S

T [Described from Russia (most sources) or Scandinavia (Fedorov 1999, Fl. Russ. 1).]



2n= 90 (2x).

2nD Löve & Löve (1975) list three counts, one Icelandic; Fl. N. Amer. 2 (1993).

G ICE RFE ALA? CAN GRL

Comments: Subsp. lanceolatum in the Arctic.


The Botrychium boreale aggregate (B. boreale, B. pinnatum)

Comments:

(1) I propose that we accept the treatment in Fl. N. Amer. 2 and separate the western North American plants as B. pinnatum. I have therefore included that species. (Elven)
5.1.4  Botrychium boreale Milde (1858), Nova Acta Phys. Med. Acad. Caes. Leop. Carol. Nat. Cur. 26, 2: 627, 757.

S

T [Described from Germany (1858?) or from Norway (1857): "drei Exemplaren stammen von Dovre, eine von Gudbrandstalen aus Norwegen".]



2n= 180 (4x).

2nD Löve & Löve (1975) list three counts, one as arctic.

G ICE NOR RUS SIB RFE GRL

Comments:

(1) Another citation   Milde (1857), Bot. Zeitung (Berlin) 15: 880   might have priority. (Elven)
5.1.5  Botrychium pinnatum H. St. John (1929), Amer. Fern J. 19: 11.

S B. boreale Milde subsp. obtusilobum (Rupr.) Clausen (1938), Mem. Torrey Bot. Club 19: 81; B. boreale auct., non Milde (1858).

2n= 180 (4x).

2nD Fl. N. Amer. 2 (1993).

G ALA CAN

Comments:


The Botrychium lunaria aggregate (B. lunaria, B. minganense)
5.1.6  Botrychium lunaria (L.) Sw. (1802), J. Bot. (Schrader) 1800, 2: 110.

B Osmunda lunaria L. (1753), Sp. Pl. 1064.

S B. lunaria (L.) Sw. subsp. lunaria.

T Clifford Herbarium 472, Osmunda 1 (BM) lectotype, selected by Jonsell & Jarvis (1994, Nordic J. Bot. 14: 148).

2n= 90 (2x).

2nD Löve & Löve (1975) list four counts, one Icelandic; Fl. N. Amer. 2 (1993).

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:


5.1.7  Botrychium minganense Vict. (1927), Proc. Trans. Roy. Soc. Canada, Ser. 3, 21: 331.

S B. lunaria (L.) Sw. var. minganense (Vict.) Dole (1937), Fl. Vermont, ed. 3, 1; B. lunaria (L.) Sw. subsp. minganense (Vict.) Calder & R.L. Taylor (1965), Canad. J. Bot. 43: 1388.

2n= 180 (4x).

2nD Fl. N. Amer. 2 (1993).

G ALA CAN GRL

Comments:

(1) Status to be discussed, as separate species or as subspecies of B. lunaria. Hultén (1968, Ark. Bot.) was very sceptical but the reported ploidy difference is an argument in favour of acceptance. At least it should be placed close to B. lunaria in the taxonomic order. It should also be looked for in material from Russian Far East. (Elven)

(2) Is the var. minganense name in use in the current standard sources? (Elven)


5.1.8  Botrychium simplex E. Hitchc. in Silliman (1823), Amer. J. Sci. Arts 6: 103, t. 8.

S

T [Described from USA: "in Conway in Massachusetts".]



2n= 90 (2x).

2nD Löve & Löve (1975) list three counts, one Icelandic; Fl. N. Amer. 2 (1993).

G GRL

Comments:



5.2  Ophioglossum L. (1753), Sp. Pl. 1062.

Comments:

(1) Tzvelev included Ophioglossum vulgatum in his draft but I have omitted it. The plants in Iceland and E Greenland, named and mapped as O. vulgatum by, e.g., Hultén & Fries (1986), are O. azoricum and were well-known as such well before 1986. The data for the species are: Ophioglossum vulgatum L. (1753), Sp. Pl. 1062. Type: Clifford Herbarium 472, Ophioglossum 1 (BM) lectotype, selected by Jonsell & Jarvis (1993, Regnum Veg. 127: 71). (Elven)
5.2.1  Ophioglossum azoricum C. Presl (1845), Suppl. Tent. Pterid. 49.

S

T [Described from the Azores.]



2n= 720 (24x).

2nD Löve & Löve (1975) list two Icelandic counts.

G GRL

Comments:



(1) The occurrences in Iceland are all outside the Arctic. There is one known arctic occurrence, in a hot spring area in E Greenland. (Elven)

6  Cryptogrammaceae (Draft: N.N. Tzvelev)

S Sinopteridaceae; Adiantaceae p. p.; Pteridaceae p. p.

Comments:

(1) Cryptogrammaceae is here proposed kept separate from Pteridaceae (and Adiantaceae). This differs from Tryon's treatment in Kubitzki (1990). (Elven)



6.1  Cryptogramma R. Br. ex Richardson in Franklin (1823), Narr. Journey Polar Sea (Bot. App.) 767.
The Cryptogramma crispa aggregate (C. acrostichoides, C. crispa, C. sitchensis)

Comments:

(1) The three taxa could be treated as species (Fl. N. Amer. 2, Tzvelev's draft) or subspecies. To be considered. (Elven)
6.1.1  Cryptogramma acrostichoides R. Br. in Franklin (1823), Narr. Journey Polar Sea (Bot. App.) 754, 767.

S C. crispa (L.) R. Br. ex Hook. var. acrostichoides (R. Br.) C.B. Clarke ***; C. crispa (L.) R. Br. ex Hook. subsp. acrostichoides (R. Br.) Hultén (1941), Lunds Univ. Årsskr., N.F., 2, 37: 40.

T [Described from Canada: Mackenzie district "in shady rocks lat. 56 north".]

2n= 60 (2x).

2nD Fl. N. Amer. 2 (1993).

G ALA CAN?

Comments:

(1) Added to Tzvelev's draft. Mapped as reaching (or at least bordering) the Arctic in Canada (Alverson 1993, Fl. N. Amer. 2) and also mapped from Nome by Hultén (1968). Should be included as species or as subspecies of C. crispa. Note, however, the reported ploidy difference between C. acrostichoides and C. crispa, which might indicate that C. crispa is an allotetraploid and that it should be kept separate from C. acrostichoides as species. (Elven)


6.1.2  Cryptogramma sitchensis (Rupr.) Moore (1857), Index Fil. 67.

B Allosorus sitchensis Rupr. (1845), Distr. Crypt. Vasc. Ross. 3: 47.

S C. crispa (L.) R. Br. ex Hook. var. sitchensis (Rupr.) C. Chr. ***.

2n= 120 (4x).

2nD Alverson in Windham (1993).

G ALA? CAN?

Comments:

(1) Border case both in Alaska (lower Yukon valley and Kuskokwim Mts) and NW Canada. Alverson (1993) refers C. sitchensis as an allopolyploid where C. acrostichoides is one of the parents, the Asian C. raddeana Fomin possibly the other. This is an argument for acceptance as a species. (Elven)



WARNING! Will be excluded if arctic occurrences not confirmed.
6.1.3  Cryptogramma crispa (L.) R. Br. ex Hook. (1842), Gen. Fil. t. 115B.

B Osmunda crispa L. (1753), Sp. Pl. 1067.

S

T Plukenet, Phytographia t. 3, f. 2 (1691) lectotype, selected by Jonsell & Jarvis (1994, Nordic J. Bot. 14: 148).



2n= 120 (8x).

2nD Löve & Löve (1975) list three counts, two as arctic; Fl. N. Amer. 2 (1993).

G ICE NOR RUS

Comments:


6.1.4  Cryptogramma stelleri (S.G. Gmel.) Prantl (1882), Bot. Jahrb. 3: 413.

B Pteris stelleri S.G. Gmel. (1768), Nov. Comment. Acad. Sci. Imp. Petrop. 12: 519.

S

T [Described from Siberia, leg. Steller?]



2n= 60 (4x).

2nD Löve & Löve (1975) list two non-arctic counts; Fl. N. Amer. 2 (1993).

G SIB? ALA

Comments:

(1) Absent from Tzvelev's draft but obviously present in the Arctic (e.g., Seward Peninsula, see Hultén 1968) and therefore added. Also occurs near the arctic boundary in NW Canada (Porcupine River area, Richardson Mts, see Cody 1996). (Elven)

7  Thelypteridaceae (Draft: N.N. Tzvelev)

7.1  Phegopteris (C. Presl) Fée (1852), Mém. Foug. 5: 242.

B Polypodium taxon Phegopteris C. Presl (1836), Tent. Pterid. 179.


7.1.1  Phegopteris connectilis (Michx.) Watt (1867), Canad. Naturalist & Quart. J. Sci., ser. 2, 3: 159.

B Polypodium connectile Michx. (1803), Fl. Bor. Amer. 2: 271.

S Dryopteris phegopteris (L.) C. Chr. (1905), Index Filic. 284; Thelypteris phegopteris (L.) Sloss. in Rydb. (1917), Fl. Rocky Mts. 1043.

T [Described from Canada.]

2n= 90 (6x).

2nD Löve & Löve (1975) list numerous counts, one Icelandic; Fl. N. Amer. 2 (1993); Jonsell (2000 Sweden).

G ICE NOR RUS RFE ALA CAN GRL

Comments:



8  Aspleniaceae (Draft: N.N. Tzvelev)

8.1  Asplenium L. (1753), Sp. Pl. 1078.
8.1.1  Asplenium viride Huds. (1762), Fl. Angl. 385.

S A. trichomanes ramosum L. (1753), nom. rejic. prop., Sp. Pl. 1082.

T [Described from Britain.]

2n= 72 (6x).

2nD Löve & Löve (1975) list numerous counts, two as arctic; Fl. N. Amer. 2 (1993).

G NOR RUS SIB ALA GRL



Comments:


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