48 Ceratophyllaceae (Draft: O. V. Rebristaya) 48. 1 Ceratophyllum

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48  Ceratophyllaceae (Draft: O.V. Rebristaya)

48.1  Ceratophyllum L. (1753), Sp. Pl. 992.
48.1.1  Ceratophyllum demersum L. (1753), Sp. Pl. 992.


2n= (1) 24. (2) 38. (3) 48.

2nD (1) Langlet & Söderberg (1927 Scand); Skalinska et al. (1978 C Eur); Váchová & Feráková (1978 C Eur); Les (1988 N Am). (2) Bir & Sidhu (1978 S As). (3) Skalinska et al. (1978 C Eur).



(1) Reaches the Arctic in Bolshezemelskaya Tundra, perhaps also in Malozemelskaya Tundra (Jalas & Suominen 1989), in the Lena River delta (Hultén & Fries 1986), and north of Porcupine River in the Yukon Territory (Cody 1996 p. 285). In Rebristaya's draft is also indicated occurrences in Russian Far East and West Alaska but the latter is not reflected in Hultén (1968) or Porsild & Cody (1980). (Elven)

(2) The deviating 2n=38 chromosome number from S Asia should be discounted. (Elven)

49  Nymphaeaceae (Draft: O.V. Rebristaya)

49.1  Nuphar Sm. in Sibth. & Sm. (1809), nom. conserv., Fl. Graec. Prodr. 1: 361.
The Nuphar lutea aggregate


(1) All three entities accepted as species below belong to the N. lutea aggregate. Nuphar polysepala and N. variegata are allopatric American vicariants of N. lutea s. str., N. pumila is a Eurasian partly sympatric northern parallel to N. lutea. A treatment as three subspecies of N. lutea is also possible. (Elven)

(2) A fourth species is given in Rebristaya's distributional table (but not in the draft) for northern Kola Peninsula - N. lutea (L.) Sm. in Sibth. & Sm. (1809), Fl. Graec. Prodr. 1: 361; basionym: Nymphaea lutea L. (1753), Sp. Pl. 510. Occurrence within the arctic parts should be confirmed before this record is accepted and the species entered in the list. On the Norwegian and Finnish side, hybrids between N. lutea and N. pumila reach much further north than N. lutea itself but they do not reach the arctic parts. (Elven)

(3) The Kola Peninsula plant is N. pumila. (Rebristaya)
49.1.1  Nuphar pumila (Timm) DC. (1821), Syst. Nat. 2: 61.

B Nymphaea lutea L. var. pumila Timm (1795), Mag. Naturk. Oekon. Meklenburgs 2: 250.


2n= 34 (2x).

2nD Löve & Löve (1975) list four non-arctic counts.



(1) Added to Rebristaya's draft. Reaches the Arctic at least in eastern parts of Bolshezemelskaya Tundra, bordering on Polar Ural (Hultén & Fries 1986, Jalas & Suominen 1989). (Elven)

49.1.2  Nuphar polysepala Engelm. (1865), Trans. Acad. Sci. St. Louis 2: 282. [Hultén 1944: p. 283]

S N. lutea (L.) Sibth. & Sm. subsp. polysepala (Engelm.) Beal ***

T [Described from Rocky Mountains.]

2n= 34 (2x).

2nD Taylor & Mulligan (1968 N Am).



(1) Endemic of northwestern North America. (Rebristaya)

(2) Reaches the Arctic as defined in the Mackenzie   Anderson Rivers area (Mackenzie District) and in N and W Alaska. (Elven)
49.1.3  Nuphar variegata Durand in Clinton (1866), Rep. (Ann.) Reg. Univ. State New York State Cab. Nat. Hist. 19: 73.


2n= 34 (2x).

2nD Löve (1954b).



(1) A border case both along Hudson Bay, near Great Bear Lake and in the Mackenzie River delta and at least approaches the Arctic in Ungava Peninsula. (Elven)

49.2  Nymphaea L. (1753), Sp. Pl. 510.
49.2.1  Nymphaea tetragona Georgi (1775), Bemerk. Reise 1: 220.

S N. alba L. subsp. tetragona (Georgi) Korsh. ***


(1) Nymphaea tetragona is the most distinct of the northern entities of the N. alba L (1753, Sp. Pl. 510) aggregate. There are arguments both for a wide, collective concept of N. alba and for a separation of N. tetragona from the rest of the complex. The latter course is followed here. (Rebristaya & Elven)  Nymphaea tetragona Georgi subsp. tetragona


2n= 112 (8x).

2nD Okada & Tamura (1981 Japan).



(1) Entered for Polar Ural in the Russian distribution table but not in Rebristaya's draft. Nymphaea tetragona s. str. also approaches the Arctic, e.g. in Alaska. (Elven)

(2) A border case in Polar Ural. (Rebristaya)

WARNING! Needs confirmation as arctic.  Nymphaea tetragona Georgi subsp. leibergii (Morong) A.E. Porsild (1939), Canad. Field-Natural. 53, 4: 50.

B N. leibergii Morong (1888), Bot. Gaz. 13: 124, in note.





(1) A border case on the SW shore of Hudson Bay (Severn River in Ontario). Wiersema (1997, Fl. N. Amer. 3) treats it as a separate species but we prefer to emphasize the relationship with N. tetragona s. str. (Rebristaya & Elven)

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