Alchemilla




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59.20  Alchemilla L. (1753), Sp. Pl. 123. [VT]

Comments:

(1) Two taxa included in Fl. Arct. URSS are excluded here: A. glaucescens Wallr., A. conglobata H. Lindb. They are also missing from Tikhomirov's draft.

There are three options for the comparatively many agamospermic 'species' in ser. Vulgares. (a) Some authors, e.g., the Löves, consider them all as subspecies of A. vulgaris L. A similar approach is used by students of some other agamospermic groups, e.g., the Ranunculus auricomus group. (b) Most authors consider the entities in Alchemilla fairly distinct as agamosperms go. It is usually not much more difficult to identify and characterise species here than in many sexual groups, with a notable exception for the A. acutidens group. This is the approach below: treatment as normal species. (c) We could also enter broader groups of entities and just list agamospecies as done below in, e.g., Antennaria, Taraxacum and Hieracium. (Elven)


Ser. Saxatiles Buser in Schroeter (1908), Pflanzenleb. Alp. 43.
59.20.1  Alchemilla alpina L. (1753), Sp. Pl. 123.

S

2n= 119-c.152



2nD Löve & Löve (1975) list six counts in this range, one Icelandic (c. 128).

G ICE NOR RUS GRL

Comments:
Ser. Splendentes Buser ***
59.20.2  Alchemilla faeroeensis (Lange) Buser (1894), Ber. Schweiz. Bot. Ges. 4: 58.

B A. alpina var. faeroeensis Lange ***

S

2n= 172-224.



2nD Löve & Löve (1975) list three counts in this range, one Icelandic (c. 224); Bozman & Walters (1967 Icel).

G ICE


Comments:

(1) Given from Jan Mayen in Tikhomirov's draft. The plant has never been reported from there but reaches the Arctic as defined in northeasternmost Iceland. (Elven)


Ser. Vulgares Buser ***
59.20.3  Alchemilla monticola Opiz in Bercht. & Opiz (1838), Oekon. Techn. Fl. Böhm. 2, 1: 13.

S A. pastoralis Buser (1892), Bull. Soc. Dauphin. Échange Pl., sér. 2, 3: 108.

2n= c. 101-109.

2nD Ehrenberg (1945 Sweden, c. 101); Turesson (1957 Norw, Sweden, 103-109).

G NOR RUS

Comments:


59.20.4  Alchemilla acutiloba Opiz in Bercht. & Opiz (1838), Oekon.-Techn. Fl. Böhm. 2, 1: 15.

S

2n= c.100-109.



2nD Ehrenberg (1945 Sweden, c.100); Lövkvist (1963 Sweden, 105-109).

G NOR* RUS*?

Comments:

(1) Alchemilla acutiloba is given in Tikhomirov's draft from Polar Ural. An occurrence is not impossible but should be checked before acceptance. If it occurs it should be clear whether it is native or introduced, stable or casual. Recently reported as stable both inside and close to arctic Norway (Alm et al. 2000). (Elven)


59.20.5  Alchemilla subcrenata Buser in Magnier (1893), Scrin. Fl. Select. 12: 285.

S A. vulgaris L. subsp. subcrenata (Buser) Murb. ***

2n= 96-c.108.

2nD Löve & Löve (1975) list four counts in this range, one (96+ff) as arctic.

G ICE NOR RUS

Comments:


59.20.6  Alchemilla filicaulis Buser (1893), Bull. Herb. Boissier 1, app. 2: 22.

S A. vulgaris L. subsp. filicaulis (Buser) Murb. (1895), Bot. Not. 1895: 265.

2n= 96-c.150.

2nD Löve & Löve (1975) list several counts in this rang, one (96+ff) as arctic.

G ICE NOR GRL

Comments:


59.20.7  Alchemilla vestita (Buser) Raunk. (1906), Dansk Exkurs. Fl., ed. 2, 145.

S A. vulgaris L. subsp. vestita (Buser) Murb. (1895), Bot. Not. 1895: 265; A. filicaulis Buser subsp. vestita (Buser) Á. Löve & D. Löve (1976), Bot. Not. 128: 515.

2n= 96-c.110.

2nD Löve & Löve (1975) list four counts in this range, one (96+ff) as arctic.

G ICE GRL

Comments:

(1) Added to Tikhomirov's draft as it reaches the Arctic both in Iceland and Greenland. This entity is often treated as a subspecies of A. filicaulis but is fairly distinct in morphology and has a more oceanic distribution. (Elven)
59.20.8  Alchemilla semispoliata Juz. (1955), Bot. Mater. Gerb. Bot. Inst. AN SSSR 17: 253.

S

2n=



2nD

G RUS


Comments:
59.20.9  Alchemilla glomerulans Buser (1893), Bull. Herb. Boissier 1, app. 2: 30.

S A. vulgaris L. subsp. glomerulans (Buser) O. Bolós & Vigo (1974), Butl. Inst. Catalana Hist. Nat. 38, Bot. 1: 68.

2n= 96-c.144.

2nD Löve & Löve (1975) list three counts in the range 96-109, two as arctic; Löve & Löve in Moore (1982 Icel, c.144).

G ICE RUS CAN GRL

Comments:  


59.20.10  Alchemilla borealis G. Sam. ex Juz. in Pojark. (1959), Fl. Murb. Obl. 4: 324, 106.

S

2n= 130-152.



2nD Turesson (1957 Norw, Sweden).

G NOR RUS

Comments:

(1) Added to Tikhomirov's draft as it reaches the Arctic in northern Norway and Kola Peninsula. (Elven)


59.20.11  Alchemilla obtusiformis Alechin in Govor. (1937), Fl. Urala 531.

S

2n=



2nD

G RUS


Comments:
59.20.12  Alchemilla transpolaris Juz. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 179.

S A. vulgaris L. subsp. transpolaris (Juz.) Á. Löve & D. Löve (1976), Bot. Not. 128: 515.

2n=

2nD


G RUS?

Comments:

(1) Not included in Tikhomirov's draft, neither as accepted species nor as synonym. Non-arctic? (Elven)

WARNING! Might be excluded as non-arctic.
59.20.13  Alchemilla baltica G. Sam. ex Juz. in Majevskii (1940), Fl. Svedi. Polosi Evrop. SSSR 7: 449.

S

2n=



2nD

G RUS


Comments:
59.20.14  Alchemilla wichurae (Buser) Stefánsson (1901), Fl. Islands 135.

S A. vulgaris L. subsp. wichurae (Buser) Gams ***

2n= 96-106.

2nD Löve & Löve (1975) list three counts in this range, one (96+ff) as arctic.

G ICE NOR RUS* GRL

Comments:


59.20.15  Alchemilla murbeckiana Buser (1906), Bot. Not. 1906: 142.

S A. vulgaris L. subsp. murbeckiana (Buser) Á. Löve ***

2n= 96-109.

2nD Löve & Löve (1975) list three counts in this range, one (96+ff) as arctic.

G NOR RUS SIB

Comments:


59.20.16  Alchemilla obtusa Buser (1894), Alchim. Valais 22.

S

2n= c. 103.



2nD Turesson (1957 Sweden).

G RUS?


Comments:

(1) Given for Polar Ural with a question mark in Tikhomirov's draft. (Elven)



WARNING! Might be excluded as non-arctic.
59.20.17  Alchemilla glabra Neygenf. (1921), Enchirid. Bot. Siles. 67.

S A. vulgaris L. subsp. alpestris (F.W. Schm.) Murb. ***; A. vulgaris L. subsp. glabra (Neyg.) O. Bolós & Vigo (1974), Butl. Inst. Catalana Hist. Nat. 38, Bot. 1: 68.

2n= 96-110.

2nD Löve & Löve (1975) list five counts in the range 96-c.104, two as arctic; Turesson (1957 Fennosc., 102-110).

G ICE NOR RUS

Comments:


59.20.18  Alchemilla cuncatrix Juz. (1951), Bot. Mater. Gerb. Bot. Inst. AN SSSR 14: 181.

S

2n=



2nD

G RUS


Comments:
59.20.19  Alchemilla glabriformis Juz. (1951), Bot. Mater. Gerb. Bot. Inst. AN SSSR 14: 178.

S

2n=



2nD

G RUS


Comments:

59.21  Sorbus L. (1753), Sp. Pl. 477. [VG]

Comments:

(1) What about Sorbus boissieri in RFE? (Elven)
59.21.1  Sorbus scopulina Greene (1899), Pittonia 4: 130.

S S. alaskana G.N. Jones (1939), J. Arnold Arb. 20: 24.

T [Described from W USA: Colorado and New Mexico.]

2n=


2nD

G ALA


Comments:

(1) More or less a border case in W Alaska, but as it approaches the border in several places (see Hultén 1968) it should be included. (Elven)


59.21.2  Sorbus aucuparia L. (1753), Sp. Pl. 477.

S

Comments:



(1) Löve & Löve (1975) also include subsp. praemorsa (Guss.) Nyman from Russian Far East. This subspecies is mentioned as Mediterranean by Flora Europaea. Opinions? (Elven)
59.21.2.1  Sorbus aucuparia L. subsp. aucuparia

S

2n= 34 (2x).



2nD ?

G NOR RUS

Comments:
59.21.2.2  Sorbus aucuparia L. subsp. glabrata (Wimm. & Grab.) Hayek (1910), Fl. Steiermark 1: 969.

B Pyrus aucuparia Ehrh. var. (b) glabrata Wimm. & Grab. (1829), Fl. Siles. 695.

S S. gorodkovii Pojark. (1966), Fl. Murm. Obl. 4: 534; S. aucuparia L. subsp. gorodkovii (Pojark.) Bondar. ex O.N. Korovina (1978), Byull. Vses. Ord. Lenina Inst. Rast. N.I. Vavilov 81: 37.

2n= 34 (2x).

2nD Löve & Löve (1975) list two Icelandic counts.

G ICE NOR RUS

Comments:
59.21.2.3  Sorbus aucuparia L. subsp. sibirica (Hedl.) Kryl. (1933), Fl. Zap. Sib. 7: 1464.

B S. sibirica Hedl. (1901), Monogr. Sorbus 44.

S S. polaris Koehne (1912), Feddes Repert. 10: 502.

2n= 34 (2x).

2nD Gladkova (1967a).

G RUS SIB

Comments:
59.21.2.4  Sorbus aucuparia L. subsp. anadyrensis (Kom.) Á. Löve & D. Löve ***

B S. anadyrensis Kom. (1921), Not. Syst. Herb. Hort. Petrop. 2: 134.

S S. sibirica auct., non Hedl. (1901) p.p.

2n=


2nD

G RFE


Comments:
59.21.3  Sorbus groenlandica (C.K. Schneid.) Á. Löve & D. Löve (1965), Univ. Colorado Stud., Biol. Ser., 17: 23.

B S. americana Marshall var. groenlandica C.K. Schneid. (1906), Bull. Herb. Boiss. 2, 6: 314.

S S. decora (Sarg.) C.K. Schneid. var. groenlandica (C.K. Schneid.) G.N. Jones (1939), J. Arnold Arb. 20: 30.

2n= 68 (4x).

2nD Böcher & Larsen (1950 Grl); Holmen in Jørgensen et al. (1958 Grl); Löve & Löve (1965, 1966b NE USA).

G GRL


Comments:

(1) What about S. decora s. str.? It is reported north to Ungava Bay by Scoggan (1978). (Elven)

(2) Also reported from 'Baffin-Labrador' in Gladkova's draft but does not seem to reach the Arctic here, acc. to Scoggan (1978) and Hultén & Fries (1986). (Elven)
59.21.4  Sorbus sambucifolia (Cham. & Schltdl.) M. Roem. (1847), Fam. Nat. Synops. 3: 139.

B Pyrus sambucifolia Cham. & Schltdl. (1827), Linnaea 2: 36.

S

T [Described from Kamtchatka: Petropavlovsk.]



2n= 34 (2x).

2nD ?


G RFE?

Comments:

(1) The records for the southern coast of Gulf of Anadyr need confirmation; non-rare in the southern part of the Korjak Mts. (Yurtsev)

WARNING! Might be excluded as non-arctic.

59.22  Amelanchier Medikus (1789), Philos. Bot. 1: 155. [VG]
59.22.1  Amelanchier alnifolia (Nutt.) Nutt. (1834), J. Acad. Nat. Sci. Philadelphia 7: 22.

B Aronia alnifolia Nutt. (1818), Gen. N. Amer. Pl. 1: 306.

S

T [Described "from Fort Mandan to the Northern Andes."]



2n=

2nD


G CAN?

Comments:

(1) The two northernmost occurrences reported by Scoggan (1978), at Gillam (Manitoba) and Fawn R. (Ontario), both seem to be south of the Arctic. Genus and species to be excluded? (Elven)

WARNING! Species and genus might be excluded as non-arctic.

59.23  Cotoneaster Medikus (1789), Philos. Bot. 1: 154. [VG]
59.23.1  Cotoneaster uniflorus Bunge in Ledeb. (1830), Fl. Alt. 220.

S C. integerrimus Medik. var. uniflorus (Bunge) Schneid. (1906), Ill. Handb. Lauholzk. 1: 747; C. integerrimus Medik. subsp. uniflorus (Bunge) Hiit. ***; C. cinnabarinus auct., non Juz. (1950).

2n= 34 (2x).

2nD Gladkova (1968).

G RUS

Comments:



(1) Does its absence from Gladkova's draft mean that C. cinnabarinus Juz. (1950), Bot. Mater. Gerb. Bot. Inst. AN SSSR 13: 32, should be excluded from the list? The northernmost occurrence in Kola Peninsula seems to be outside the Arctic as defined. (Elven)
59.23.2  Cotoneaster niger (Wahlb.) Fr. (1846), Summa Veg. Scand. 1: 175.

B Mespilus cotoneaster L. var. (beta) nigra Wahlb. (1820), Fl. Gothob. 1: 53.

S C. melanocarpus auct., non Loddiges (1829).

T Sweden: På näset vid Varberg, 1819, leg. Wahlberg (UPS, ex herb. Wahlenberg) lectotype, selected by Hylmö (1993, Sv. Bot. Tidskr. 87: 315).

2n= 68 (4x).

2nD Löve & Löve (1975) list three non-arctic counts.

G RUS

Comments:



(1) N Fennoscandia added as it is mapped north of Ponoj by Fl. Murm. Obl. However, Fl. Arct. URSS does not report it from the Kanin -Pechora area where it is given in Gladkova's draft. (Elven)

(2) Cotoneaster melanocarpus is described on garden material, possibly of S Russian origin and not certainly identical with the wild plant often named as such. It is also some doubts concerning the description, whether it is valid. (Elven)



59.24  Prunus L. (1753), Sp. Pl. 473. [VG]

S Padus Mill. (1754), Gard. Dict. Abr., ed. 4.

Comments:

(1) Genus concept must be discussed, whether to separate Padus from Prunus s. str. or not. As this separation is not usually applied in W Europe and North America, I here tentatively have changed Gladkova's Padus to Prunus but I can see arguments for both solutions. (Elven)

(2) Only P. padus subsp. padus is included in the draft received from Gladkova. Some part may be missing. The 'borealis' entity must probably appear somewhere, at least in synonymy, but I am not sure that it reaches the Arctic. I have tentatively entered it as a subspecies. Prunus pensylvanica is also entered as a border case. (Elven)
59.24.1  Prunus padus L. (1753), Sp. Pl. 473.

S Padus avium Mill. (1768), Gard. Dict., ed. 8, no. 1.

Comments:
59.24.1.1  Prunus padus L. subsp. padus

S Padus avium Mill. subsp. avium

2n= 32 (4x).

2nD Lövkvist (1963 Sweden).

G NOR? RUS

Comments:

(1) There are no specimens in HbO from the arctic parts of Norway. I will check with TROM. (Elven)
59.24.1.2  Prunus padus L. subsp. borealis (Schuebeler) Nyman (1878), Consp. Fl. Eur. 212.

B P. borealis Schuebeler (1873-1875), Pflanzenw. Norw. 369.

S Padus avium Mill. subsp. borealis (Schuebeler) Holub (1970), Preslia 42, 1: 94.

T Two specimens available for typification, both in HbO: (1) "Prunus borealis Sch.", Schübeler scripsit, probably from Alta; (2) "Prunus padus L. var. ?", Alten, 1870, leg. Schübeler (scripsit).

2n= 32 (4x).

2nD Knaben & Engelskjøn (1967 Norw).

G RUS?

Comments:



(1) The occurrences mapped in European Russia in Fl. Arct. URSS all seem to be south of the Arctic as defined. There are no specimens from the arctic parts of Norway in HbO; I will check with TROM. (Elven)

WARNING! Might be excluded as non-arctic.
59.24.2  Prunus pensylvanica L. fil. (1782), Suppl. Pl. 252.

S

2n=



2nD

G CAN


Comments:

(1) A border case on the S and SE shores of Hudson Bay, see Scoggan (1978) and Porsild & Cody (1980): Manitoba: Cochrane R. (5813'N), Ontario: Fawn R. (54N), Quebec: Swampy R. (5606'N). (Elven)


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